Dianthus (Velezia rigida) 1 ESE Filia, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus (Velezia rigida) 2 ESE Filia, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 2b Dadia, Thracia, Greece 060603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 3 3 km SW of Polygyros, Greece 050526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 5a Lund Bot Garden 160609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 5a Parque Nacional de Monfragüe SP 140507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 5b Lund Bot Garden 160609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 5b Parque Nacional de Monfragüe SP 140507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 5c Parque Nacional de Monfragüe SP 140507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 5d Parque Nacional de Monfragüe SP 140507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 6a near Trujillo SP 140507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 6b near Trujillo SP 140507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 7a Skurup Sk 140914 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 7b Skurup Sk 140914 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 8a Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 8b Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus aff cruentus 1 W of Mandra, Thracia, Greece 060605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus aff cruentus 2 W of Mandra, Thracia, Greece 060605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus alpinus 2 Lund Bot Garden 130610 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus alpinus 3 Lund Bot Garden 130610 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus amurensis 2 Lund Bot Garden 120907 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus amurensis 3 Copenhagen Bot Garden 130807 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus andrzejowskianus 2 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arenarius ssp arenarius 1 Vitemölla, Vitaby Sk 120609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arenarius ssp arenarius 2 between Kivik and Vitemölla, Vitaby Sk 080603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arenarius ssp arenarius 3 Vitemölla, Vitaby Sk 120609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arenarius ssp arenarius 4 Vitemölla, Vitaby Sk 220609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arenarius ssp arenarius 5 Vitemölla, Vitaby Sk 220609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arenarius ssp borussicus 1 Kehgarags in Riga, Latvia 060711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arenarius ssp borussicus 2 Kehgarags in Riga, Latvia 060711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arenarius ssp borussicus 3 main road near road to Saaramaa, Valkeala FI 120730 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arenarius ssp borussicus 4 main road near road to Saaramaa, Valkeala FI 120730 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arenarius ssp borussicus 5 main road near road to Saaramaa, Valkeala FI 120730 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 10 (cult) Brunnby Sk 140717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 11 (cult) Brunnby Sk 140717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 2 Sölve grustag, Sölvesborg, Blekinge 930815 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 7 (cult) Brunnby Sk 140717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 8 (cult) Brunnby Sk 140717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 9 (cult) Brunnby Sk 140717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arpadianus 1 Lund Bot Garden 160615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arpadianus 2 Lund Bot Garden 160615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arpadianus 3 Lund Bot Garden 160615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus arpadianus 4 Lund Bot Garden 160615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus barbatus 2 Lund Sk 130627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus barbatus 3 Lund Sk 130627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus barbatus x superbus 1 Klagshamn, Skåne 780812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus barbatus x superbus 2 Klagshamn, Skåne 780812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus bisignani Bergianska, Stockholm 050621 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 10 near Merkine, Lithuania 150806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 3 Merkine, Lithuania 100714 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 6 near Merkine, Lithuania 150806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 7 near Merkine, Lithuania 150806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 8 near Merkine, Lithuania 150806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 9 near Merkine, Lithuania 150806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus broteri 1 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus broteri 2 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus broteri 3 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus broteri 4 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum 1 Copenhagen Bot Garden 120926 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum 2 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum 4 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum 5 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum 6 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum 7 Hosszúhetény, Nagy-mezö, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum 8 Hosszúhetény, Nagy-mezö, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum 9 Hosszúhetény, Nagy-mezö, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum ssp atrorubens 2 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum ssp atrorubens 3 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum ssp atrorubens 4 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum ssp atrorubens 5 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus caryophyllus 1 Lund Bot Garden 130619 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus caryophyllus 2 Lund Bot Garden 090708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cf glutinosus 1 Sigri, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cf glutinosus 2 Mikri Limni, Lesvos, Greece 100511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cf glutinosus 3 Mikri Limni, Lesvos, Greece 100511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cf glutinosus 4 Mikri Limni, Lesvos, Greece 100511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cf petraeus 1 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cf petraeus 2 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cf pungens 1 Lund Bot Garden 150713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cf pungens 2 Lund Bot Garden 150713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 1 Rommunds, Gammelgarn, Gotland 830627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 10 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 11 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 12 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 13 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 3 Rommunds, Gammelgarn, Gotland 830627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 5 Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 6 Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 7 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 8 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 9 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus deltoides 2 Höllviken Sk 120623 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus furcatus 1 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis 6 Malmö Sk 200723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 10 Hosszúhetény, Nagy-mezö, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 2 Lund Bot Garden 130619 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 3 Lund Bot Garden 130619 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 6 Nagykovácsi, Hungary 240504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 7 Nagykovácsi, Hungary 240504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 8 Nagykovácsi, Hungary 240504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 9 Hosszúhetény, Nagy-mezö, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteus 1 Lund Sk 200716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteus 2 Lund Sk 200716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteus 3 Lund Sk 200716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteus 4 Lund Sk 200716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteus 5 Lund Sk 200716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus haematocalyx ssp pindicola 1 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus haematocalyx ssp pindicola 2 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus haematocalyx ssp pindicola 3 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus haematocalyx ssp pindicola 4 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus haematocalyx ssp pindicola 5 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus haematocalyx ssp pindicola 6 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus haematocalyx ssp pindicola 7 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus knappii (cult) Stångby, Skåne 050717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus knappii 1 Lund Bot Garden 160627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus knappii 4 Lund Bot Garden 160627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus knappii 5 Lund Bot Garden 160627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus knappii 6 Lund Bot Garden 160627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus lumnitzeri 1 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus lumnitzeri 2 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus monspessulanus 1 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus monspessulanus 3 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus monspessulanus 4 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus monspessulanus 5 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus monspessulanus 6 Mont Aigoual, Cévennes Nat. Park, France 180720 - Photo: Ragnhild Crawford (© 2014 Jan Thomas Johansson).

Dianthus myrtinervius 1 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus myrtinervius 2 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus myrtinervius 3 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pavoninus 1 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pavoninus 2 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus petraeus 1 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus petraeus 3 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus petraeus 4 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus petraeus ssp petraeus 1 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus petraeus ssp petraeus 2 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pinifolius 1 Monis Gorgianis near old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pinifolius 2 Monis Gorgianis near old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pinifolius 4 Monis Gorgianis near old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pinifolius 5 Monis Gorgianis near old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus plumarius 3 Lund Bot Garden 130612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus plumarius 4 Lund Bot Garden 130612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus plumarius ssp regis-stephani 1 Nagykovácsi, Hungary 240504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus plumarius ssp regis-stephani 2 Nagykovácsi, Hungary 240504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pygmaeus 1 Lund Bot Garden 150731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pygmaeus 2 Lund Bot Garden 150713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pyrenaicus 1 Lund Bot Garden 080612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pyrenaicus 3 Lund Bot Garden 080612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus sanguineus 1 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus sanguineus 2 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 1 Comboire FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 10 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 11 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 12 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 2 Comboire FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 3 between Lans en Vercors and Villard-de-Lans FR 150702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 4 between Lans en Vercors and Villard-de-Lans FR 150702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 5 between Lans en Vercors and Villard-de-Lans FR 150702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 7 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 8 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 9 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus seguieri 2 Lund Bot Garden 150925 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus seguieri 3 Lund Bot Garden 150925 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus seguieri 4 Lund Bot Garden 150925 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus seguieri 5 Lund Bot Garden 190720 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus seguieri 6 Lund Bot Garden 190720 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus seguieri 7 Lund Bot Garden 190720 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus seguieri 8 Lund Bot Garden 190720 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus serotinus 2 Lund Bot Garden 150713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus spiculifolius 1 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus spiculifolius 3 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus squarrosus 1 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus squarrosus 2 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus sternbergii 1 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus sternbergii 2 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus sternbergii 3 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus sternbergii 6 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 1 NE of Hårbölle, Mön DK 080824 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 11 near Båtsfjord, between Hamningberg and Vardø, Finnmark, Norway 220725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 12 near Båtsfjord, between Hamningberg and Vardø, Finnmark, Norway 220725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 14 near Båtsfjord, between Hamningberg and Vardø, Finnmark, Norway 220725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 15 near Båtsfjord, between Hamningberg and Vardø, Finnmark, Norway 220725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 16 near Båtsfjord, between Hamningberg and Vardø, Finnmark, Norway 220725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 17 near Båtsfjord, between Hamningberg and Vardø, Finnmark, Norway 220725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 2 NE of Hårbölle, Mön DK 080824 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 3 NE of Hårbölle, Mön DK 080824 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 4 NE of Hårbölle, Mön DK 080824 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 5 Lilla Hammar, Stora Hammar Sk 190724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 6 Lilla Hammar, Stora Hammar Sk 190724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 7 Lilla Hammar, Stora Hammar Sk 190724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 9 Lilla Hammar, Stora Hammar Sk 190724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus tripunctatus 1 Kampos, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus tripunctatus 2 Kampos, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus turkestanicus 2 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus turkestanicus 3 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus turkestanicus 4 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus turkestanicus 5 Lund Bot Garden 130610 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus turkestanicus 6 Lund Bot Garden 130610 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus versicolor 1 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus versicolor 2 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus versicolor 3 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus viscidus 2 Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus viscidus 4 Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus viscidus 5 Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus viscidus 6 Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus viscidus 9 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus waldsteinii 1 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus waldsteinii 2 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 2a Dadia, Thracia, Greece 060603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus 5c Lund Bot Garden 160609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus aff cruentus 3 Holomondas, Greece 050528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus alpinus 1 Lund Bot Garden 130610 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus amurensis 1 Copenhagen Bot Garden 130807 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus anatolicus 1 Lesvos, Greece 100511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus anatolicus 2 Lesvos, Greece 100511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus anatolicus 3 Lesvos, Greece 100511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus anatolicus 4 Lesvos, Greece 100511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus andrzejowskianus 1 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 1 Munka-Ljungby, Skåne 990723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 3 (cult) Brunnby Sk 140717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 4 (cult) Brunnby Sk 140717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 5 (cult) Brunnby Sk 140717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus armeria 6 (cult) Brunnby Sk 140717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus barbatus 1 Lund Sk 130627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 1 Merkine, Lithuania 100714 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 11 near Merkine, Lithuania 150806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 2 Merkine, Lithuania 100714 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 4 Merkine, Lithuania 100714 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus borbasii 5 Merkine, Lithuania 100714 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus caespitosus 1 Little Karoo Desert National Bot Garden, Worcester, Western Cape, South Africa 161005 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus caespitosus 2 Little Karoo Desert National Bot Garden, Worcester, Western Cape, South Africa 161005 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus caespitosus 3 Little Karoo Desert National Bot Garden, Worcester, Western Cape, South Africa 161005 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum 3 Copenhagen Bot Garden 120926 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus carthusianorum ssp atrorubens 1 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 2 Rommunds, Gammelgarn, Gotland 830627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus cruentus 4 Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus deltoides 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus deltoides 3 Lund Sk 200721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus deltoides 4 Lund Sk 200721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus furcatus 2 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis 2 Malmö Sk 200723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis 3 Malmö Sk 200723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis 4 Malmö Sk 200723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis 5 Malmö Sk 200723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis 7 Norra hamnen, Malmö Sk 200919 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis 8 Norra hamnen, Malmö Sk 200919 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 1 Lund Bot Garden 130619 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 11 Hosszúhetény, Nagy-mezö, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 4 Nagykovácsi, Hungary 240504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus giganteiformis ssp pontederae 5 Nagykovácsi, Hungary 240504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus haematocalyx ssp haematocalyx 1 near Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus knappii 2 Lund Bot Garden 160627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus knappii 3 Lund Bot Garden 160627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus monspessulanus 2 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus nitidus W of Zilina, Czekia 950630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus petraeus 2 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus petraeus ssp orbelicus 1 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus petraeus ssp orbelicus 2 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus petraeus ssp orbelicus 3 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus petraeus ssp orbelicus 4 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pinifolius 3 Monis Gorgianis near old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus plumarius 1 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus plumarius 2 Lund Bot Garden 130612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pygmaeus 3 Lund Bot Garden 150731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pygmaeus 4 Lund Bot Garden 150731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pygmaeus 5 Lund Bot Garden 150731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pygmaeus 6 Lund Bot Garden 150713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus pyrenaicus 2 Lund Bot Garden 080612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 13 Les Freaux near La Meije FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 14 Les Freaux near La Meije FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 15 Les Freaux near La Meije FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus saxicola 6 between Lans en Vercors and Villard-de-Lans FR 150702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus seguieri 1 Lund Bot Garden 150925 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus serotinus 1 Lund Bot Garden 150713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus spiculifolius 2 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus sternbergii 4 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus sternbergii 5 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus sternbergii 7 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 10 Lilla Hammar, Stora Hammar Sk 190724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus superbus 8 Lilla Hammar, Stora Hammar Sk 190724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus tripunctatus 3 Kampos, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus tripunctatus 4 Kampos, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus turkestanicus 1 Lund Bot Garden 130610 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus viscidus 1 Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus viscidus 3 Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus viscidus 7 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Dianthus viscidus 8 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).