Berberis 1a Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 1b Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 1c Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 1d Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 1e Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 2a Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 2b Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 3a Skanör Sk 140417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 3b Skanör Sk 140417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 3c Skanör Sk 140417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 3d Skanör Sk 140417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 3e Skanör Sk 140417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 4a Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 4b Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 5a (cult) Lund Sk 170514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 5b (cult) Lund Sk 170514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 5c (cult) Lund Sk 170514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 5d (cult) Lund Sk 170514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 5e (cult) Lund Sk 170514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 6a (cult) 170522 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 6b (cult) 170522 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 6d (cult) 170522 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 6f (cult) 170522 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 7a Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 7b Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis angulosa 1 Lund Bot Garden 080519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis angulosa 2 Lund Bot Garden 080519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis angulosa 3 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis angulosa 4 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis bealei 1 Copenhagen Bot Garden 150421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis bealei 4 Copenhagen Bot Garden 150421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis buxifolia 1 Copenhagen Bot Garden 150421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis buxifolia 2 Copenhagen Bot Garden 150421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 1 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 10 Lund Bot Garden 150520 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 11 Lund Bot Garden 150520 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 12 Lund Bot Garden 150520 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 13 Lund Bot Garden 150520 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 2 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 3 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 4 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 5 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 6 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 7 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 8 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis candidula 9 Lund Bot Garden 150520 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis cf montana 1 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis cf montana 2 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis cf montana 3 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis cf montana 4 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis cf montana 5 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis cf montana 6 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis circumserrata 1 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis circumserrata 2 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis consimilis 1 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis consimilis 2 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis consimilis 3 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis declinata 1 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis declinata 2 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis declinata 3 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis declinata 4 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis dictyophylla 1 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis dictyophylla 2 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis dielsiana 2 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis dielsiana 3 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis faxeniana 1 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis faxeniana 2 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis faxeniana 3 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis faxeniana 4 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis faxeniana 5 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis faxeniana 6 Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis faxeniana 7 Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis gagnepainii 1 Lund Bot Garden 130612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis gagnepainii 2 Lund Bot Garden 130612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis gagnepainii 3 Lund Bot Garden 130612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis gagnepainii 4 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis gagnepainii 5 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis gagnepainii 6 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis gagnepainii 7 Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis henryana 1 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis henryana 3 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis kansuensis 1 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis kansuensis 2 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis koreana 1 Copenhagen Bot Garden 130913 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis koreana 2 Copenhagen Bot Garden 130913 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis lycium 1 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis lycium 2 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis mahonia 1 (cult) Höllviken Sk 080503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis mahonia 2 (cult) Höllviken Sk 080503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis mahonia 3 (cult) Höllviken Sk 060811 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis mahonia 4 (cult) Höllviken Sk 060811 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis mahonia 5 (cult) Höllviken Sk 060811 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis muliensis var atuntzeana 1 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis muliensis var atuntzeana 2 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis repens 1 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis repens 2 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis repens 5 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis repens 6 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis repens 7 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis sibirica 1 Lund Bot Garden 130603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis sibirica 2 Lund Bot Garden 130603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis sibirica 3 Lund Bot Garden 130603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis sibirica 4 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis tsarongensis 1 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis tsarongensis 2 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis tsarongensis 3 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis turcomanica 2 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis turcomanica 3 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis turcomanica var integerrima 2 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vernae 1 Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vernae 2 Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vernae 3 Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vulgaris 1 Lund Bot Garden 130603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vulgaris 2 Lund Bot Garden 130603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vulgaris 3 Lund Bot Garden 130603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vulgaris 4 Lund Bot Garden 130603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vulgaris 5 Lund Bot Garden 130603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vulgaris 6 (cult) Bot Dept, Stockholm Univ 030909 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 6c (cult) 170522 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis 6e (cult) 170522 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis bealei 2 Copenhagen Bot Garden 150421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis bealei 3 Copenhagen Bot Garden 150421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis cf montana 7 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis declinata 5 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis dielsiana 1 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis gagnepainii 10 Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis gagnepainii 8 Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis gagnepainii 9 Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis henryana 2 Lund Bot Garden 140521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis kansuensis 3 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis kansuensis 4 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis kansuensis 5 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis kansuensis 6 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis koreana 3 Copenhagen Bot Garden 130913 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis lycium 1 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis lycium 2 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis lycium 3 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis repens 3 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis repens 4 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis turcomanica 1 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis turcomanica var integerrima 1 Lund Bot Garden 121016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vernae 4 Lund Bot Garden 141011 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vulgaris 7 Lund Bot Garden 130919 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Berberis vulgaris 8 Lund Bot Garden 130919 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).