Aquilegia 1a Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 1b Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 1c Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 1d Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 1e Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 1f Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 3b Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 3c Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia alpina 1 (cult) 190519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia alpina 2 (cult) 190519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia alpina 3 (cult) 190519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia alpina 4 (cult) 190519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia alpina 5 (cult) 190519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia alpina 6 (cult) 190519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia alpina 7 (cult) 190519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia alpina 8 (cult) 190519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia alpina 9 (cult) 190519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia artensis 1 Göteborg Bot Garden 130524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia artensis 2 Göteborg Bot Garden 130524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia atrata 2 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia atrata 3 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia canadensis 2 Flinck's Garden, Bjuv Sk 140518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia cf chrysantha 1 Desert Botanical Garden, Phoenix, Arizona 130426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia cf chrysantha 10 Desert Botanical Garden, Phoenix, Arizona 130426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia cf chrysantha 2 Desert Botanical Garden, Phoenix, Arizona 130426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia cf chrysantha 3 Desert Botanical Garden, Phoenix, Arizona 130426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia cf chrysantha 4 Desert Botanical Garden, Phoenix, Arizona 130426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia cf chrysantha 5 Desert Botanical Garden, Phoenix, Arizona 130426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia cf chrysantha 7 Desert Botanical Garden, Phoenix, Arizona 130426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia ecalcarata 2 Lund Bot Garden 130528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia einseleana 1 Lund Bot Garden 140524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia einseleana 2 Lund Bot Garden 140524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia einseleana 5 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia flabellata 1 Göteborg Bot Garden 130524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia flabellata 2 Göteborg Bot Garden 130524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia flabellata 3 Göteborg Bot Garden 130524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia formosa 2 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia formosa 3 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia formosa 4 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia formosa 5 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia formosa 6 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia hirsutissima 1 San Juan de la Peña, the Belchite area, NE Spain 180523 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia hirsutissima 2 San Juan de la Peña, the Belchite area, NE Spain 180523 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia hirsutissima 3 San Juan de la Peña, the Belchite area, NE Spain 180523 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia hirsutissima 4 San Juan de la Peña, the Belchite area, NE Spain 180523 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia kareliniana 2 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia kareliniana 3 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia kareliniana 4 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia kareliniana 5 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia oxysepala 3 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia oxysepala 4 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia oxysepala 5 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia pyrenaica 3 Lund Bot Garden 140524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia pyrenaica 4 Lund Bot Garden 140524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia rockii 2 Napa Hai, Yunnan 110709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 1 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 2 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 9 Lund Bot Garden 110429 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viscosa 3 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia vulgaris 4 near Kungsmarken, Södra Sandby Sk 140516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia vulgaris 5 near Kungsmarken, Södra Sandby Sk 140516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia vulgaris 6 near Kungsmarken, Södra Sandby Sk 140516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 2a Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 2b Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 2c Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 2d Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia 3a Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia atrata 1 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia canadensis 1 Storrs, Connecticut 900523 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia canadensis 3 Flinck's Garden, Bjuv Sk 140518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia canadensis 4 Lund Bot Garden 100527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia canadensis 5 Lund Bot Garden 100527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia canadensis 6 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia canadensis 7 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia cf chrysantha 6 Desert Botanical Garden, Phoenix, Arizona 130426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia ecalcarata 1 Lund Bot Garden 130528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia einseleana 3 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia einseleana 4 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia formosa 1 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia formosa 7 Lund Bot Garden 160615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia formosa 8 Lund Bot Garden 160615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia kareliniana 1 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia nigrescens 1 Göteborg Bot Garden 130524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia nigrescens 2 Göteborg Bot Garden 130524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia oxysepala 1 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia oxysepala 2 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia pyrenaica 2 Lund Bot Garden 140524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia rockii 1 Napa Hai, Yunnan 110709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia rockii 3 Napa Hai, Yunnan 110709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia rockii 4 Napa Hai, Yunnan 110709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 10 Göteborg Bot Garden 130524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 11 Göteborg Bot Garden 130524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 3 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 4 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 5 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 6 Lund Bot Garden 110429 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 7 Lund Bot Garden 110429 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viridiflora 8 Lund Bot Garden 110429 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viscosa 1 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia viscosa 2 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia vulgaris 1 Blombergs Säteri, Husaby, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia vulgaris 2 Blombergs Säteri, Husaby, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Aquilegia vulgaris 3 near Kungsmarken, Södra Sandby Sk 140516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).