Cynoglossum 1d Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum amabile 2 between Dali and Lijiang, Yunnan 110706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum amabile 3 between Dali and Lijiang, Yunnan 110706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum amabile 4 between Dali and Lijiang, Yunnan 110706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cf pustulatum 3 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cf pustulatum 4 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 1 El Torcal de Antequera Andulusia SP 120323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 10 Pantallica area, Sicily 190416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 12 Pantallica area, Sicily 190416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 13 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 14 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 15 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 16 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 17 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 19 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 2 El Torcal de Antequera Andulusia SP 120323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 20 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 21 Desfiladero de Los Gaitanes, Andalucía Spain 220411 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 22 Desfiladero de Los Gaitanes, Andalucía Spain 220411 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 23 Desfiladero de Los Gaitanes, Andalucía Spain 220411 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 4 Miradór del Guarda Forestal ESE Ronda, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 7 Antequera Andulusia SP 120323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 8 Pantallica area, Sicily 190416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 9 Pantallica area, Sicily 190416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum columnae 1 N Agia Galini, Crete 110329 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum columnae 2 N Agia Galini, Crete 110329 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum columnae 3 N Agia Galini, Crete 110329 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 1 Alanya, Turkey 090405 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 10 Santa Cilia, N Spain 180527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 11 Río de la Miel, Los Alcornocales Natural Park, Andalucía Spain 220413 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 12 Río de la Miel, Los Alcornocales Natural Park, Andalucía Spain 220413 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 2 Alanya, Turkey 090405 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 3 Myrsini-Sfaki, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 8 Barr. de las Cuevas Negras, Tenerife 160409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 9 Santa Cilia, N Spain 180527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum glochidiatum 2 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum glochidiatum 3 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum montanum 5 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum montanum 7 Mórahalom, Hungary 240509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum nervosum 5 Lund Bot Garden 080926 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 10 Maglarp Sk 150620 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 4 Kåseberga, Valleberga, Sk 120526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 5 Hammarbyhöjden, Stockholm 050822 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 9 Maglarp Sk 150620 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 1 Yeghegnadzan mountains, Armenia 180513 - Photo: Kim Frost (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 13 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 14 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 17 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 18 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 19 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 20 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 6 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 7 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum umbellatum 2 Lund Bot Garden 150514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum umbellatum 3 Lund Bot Garden 150514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum umbellatum 4 Lund Bot Garden 150514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum umbellatum 5 Lund Bot Garden 150514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum wallichii 1 Copenhagen Bot Garden 130913 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum wallichii 2 Copenhagen Bot Garden 130913 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum 1a Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum 1b Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum 1c Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum amabile 1 between Dali and Lijiang, Yunnan 110706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum caespitosum Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cf pustulatum 1 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cf pustulatum 2 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 11 Pantallica area, Sicily 190416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 18 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 3 El Torcal de Antequera Andulusia SP 120323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 5 Antequera Andulusia SP 120323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum cheirifolium 6 Antequera Andulusia SP 120323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum columnae 4 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 4 Barr. de las Cuevas Negras, Tenerife 160409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 5 Barr. de las Cuevas Negras, Tenerife 160409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 6 Barr. de las Cuevas Negras, Tenerife 160409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum creticum 7 Barr. de las Cuevas Negras, Tenerife 160409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum furcatum 1 Yunnan 201107 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum furcatum 2 Yunnan 201107 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum glochidiatum 1 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum hungaricum 1 NW of Straznice, Czekia 950701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum hungaricum 2 NW of Straznice, Czekia 950701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum montanum 1 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum montanum 3 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum montanum 4 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum montanum 6 Mórahalom, Hungary 240509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum montanum 8 Mórahalom, Hungary 240509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum nervosum 1 Lund Bot Garden 100701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum nervosum 2 Lund Bot Garden 080926 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum nervosum 3 Lund Bot Garden 070524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum nervosum 4 Lund Bot Garden 090819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 1 Kåseberga, Valleberga, Sk 120526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 11 Maglarp Sk 150620 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 12 Busene, Mön DK 160529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 2 Kåseberga, Valleberga, Sk 120526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 3 Kåseberga, Valleberga, Sk 120526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 6 Limhamn Sk 150625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 7 Limhamn Sk 150625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum officinale 8 Limhamn Sk 150625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 10 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 11 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 12 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 15 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 16 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 2 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 4 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 5 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 8 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum stamineum 9 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cynoglossum umbellatum 1 Lund Bot Garden 150514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).