Carduus acanthoides 4 Skanör Sk 130809 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus acicularis 3 Eleas, Lesvos 130507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus acicularis 4 Eleas, Lesvos 130507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus acicularis 5 Eleas, Lesvos 130507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus acicularis 6 Eleas, Lesvos 130507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus baeocephalus 3 Anden Verde, Gran Canaria 080412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf kerneri 2 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf kerneri 3 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf kerneri 4 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf kerneri 5 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf nutans 2 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf tenuiflorus 4 Magaluf, Mallorca 110422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf tenuiflorus 5 Magaluf, Mallorca 110422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus clavulatus 2 Teno, Tenerife 090513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus clavulatus 3 Teno, Tenerife 090513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus clavulatus 4 Teno, Tenerife 090513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus clavulatus 5 Teno, Tenerife 160404 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus crispus 3 Lund Sk 121001 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus defloratus 1 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus defloratus 2 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus defloratus 3 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus defloratus 6 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus defloratus 7 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus defloratus 8 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus defloratus 9 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus glaucus 3 Garda de Sus, Romania 090615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus glaucus 4 Garda de Sus, Romania 090615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus macrocephalus 1 above Pefkochori, Greece 050527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 11 Ar, Fleringe, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 12 Ar, Fleringe, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 13 Ar, Fleringe, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 14 Ar, Fleringe, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 2 NW of Hofterup, Skåne 790627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans ssp nutans 11 N of La Vachette FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans ssp nutans 12 N of La Vachette FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans ssp nutans 13 N of La Vachette FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus personata 3 Bad Gastein, Austria 080704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus personata 4 Garda de Sus, Romania 090615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus pycnocephalus 1 Pefkochori, Greece 050524 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus pycnocephalus 10 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus pycnocephalus 2 Lund Bot Garden 130702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus pycnocephalus 4 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus pycnocephalus 6 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus pycnocephalus 7 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus pycnocephalus 8 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus pycnocephalus 9 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus tenuiflorus 2 near Trujillo SP 140507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus tenuiflorus 3 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus tenuiflorus 4 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus tenuiflorus 5 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus tenuiflorus 6 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus acanthoides 1 Höllviken Sk 120812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus acanthoides 2 Höllviken Sk 120721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus acanthoides 3 Höllviken Sk 120721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus acicularis 1 Eleas, Lesvos 130507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus acicularis 2 Eleas, Lesvos 130507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus baeocephalus 1 Anden Verde, Gran Canaria 080410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus baeocephalus 2 Anden Verde, Gran Canaria 080412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf acanthoides 1 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf kerneri 1 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf kerneri 6 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf nutans 1 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf tenuiflorus 1 Magaluf, Mallorca 110422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf tenuiflorus 2 Magaluf, Mallorca 110422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus cf tenuiflorus 3 Magaluf, Mallorca 110422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus clavulatus 1 Pista de los Lomos, La Palma 090427 (Richard Åkesson) - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus clavulatus 6 Teno, Tenerife 160404 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus crispus 1 Lund Sk 121001 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus crispus 2 Skåne-Tranås Sk 070706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus defloratus 4 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus defloratus 5 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus glaucus 1 Garda de Sus, Romania 090615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus glaucus 2 Garda de Sus, Romania 090615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus macrocephalus 2 N of Gerakini, Greece 050525 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 1 Kyllaj, Gotland 980725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 10 Ar, Fleringe, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 3 Ljugarn, Gotland 730707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 4 Dämba, Fårö, Gotland 040703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 5 Dämba, Fårö, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 6 Dämba, Fårö, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 7 Dämba, Fårö, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 8 Ar, Fleringe, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans 9 Ar, Fleringe, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans ssp nutans 10 N of La Vachette FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans ssp nutans 8 Les Freaux near La Meije FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus nutans ssp nutans 9 Les Freaux near La Meije FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus personata 1 Garda de Sus, Romania 090615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus personata 2 Bad Gastein, Austria 080704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus pycnocephalus 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus pycnocephalus 5 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus tenuiflorus 1 near Trujillo SP 140507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus thoermeri 1 Toftemo, Norway 820801 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus thoermeri 2 Toftemo, Norway 820801 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Carduus thoermeri 3 Toftemo, Norway 820801 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).