Centaurea 2a Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea 2b Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea 2c Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea adamii 1 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea adamii 2 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea adamii 3 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea adamii 5 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea adamii 6 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea affinis 3 near Milea, old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea americana 1 Lund Bot Garden 080905 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea americana 2 Lund Bot Garden 080905 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea bella 1 Horsabäck Garden, Våxtorp Hl 130617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea bella 2 Horsabäck Garden, Våxtorp Hl 130617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea bella 3 Horsabäck Garden, Våxtorp Hl 130617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea benedictus 1 to Konya, NW of Alanya, Turkey 090409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea benedictus 2 to Konya, NW of Alanya, Turkey 090409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea benedictus 3 to Konya, NW of Alanya, Turkey 090409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea benedictus 4 above Pefkochori, Greece 050527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea bourgaei 1 SE of Megali Limni, Lesbos, Greece 100515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea bourgaei 2 SE of Megali Limni, Lesbos, Greece 100515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea calocephala 3 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea calocephala 4 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf affinis 2 Paranesti, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf alba 1 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf alba 2 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf aspera 1 Jardin Botanico, Soller, Mallorca 110419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf aspera 2 Jardin Botanico, Soller, Mallorca 110419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf aspera 3 Magaluf, Mallorca 110422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf aspera 4 Magaluf, Mallorca 110422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf aspera 5 Magaluf, Mallorca 110422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf aspera 6 Magaluf, Mallorca 110422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf nervosa ssp promota 2 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf nervosa ssp promota 3 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf nervosa ssp promota 4 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea clementei 2 Sierra del Co, Villanueva del Cauche, Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea clementei 3 Sierra del Co, Villanueva del Cauche, Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea collina 1 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea collina 2 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea collina 3 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cyanus 2 Dalby Västermark Sk 130618 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cyanus 3 Dalby Västermark Sk 130618 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea diffusa 1 Tallinn, Estonia 100708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea diluta 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea eriophora 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea eriophora 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea eriophora 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea fischeri 1 (cult) Brunnby Sk (orig Tingstäde, Gotland) 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea fischeri 2 (cult) Brunnby Sk (orig Tingstäde, Gotland) 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea fischeri 2 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea fischeri 3 Tingstäde, Gotland 980728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea fischeri 3 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea fischeri 4 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea glastifolia 1 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea glastifolia 2 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea glastifolia 3 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 10 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 11 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 12 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 13 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 14 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 15 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 2 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 6 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 7 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 8 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 9 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea horrida 1 Asinara, Sardinia, Italy 230420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea horrida 2 Asinara, Sardinia, Italy 230420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea involucrata 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea involucrata 1 Ljunghusen, Stora Hammar Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea involucrata 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea involucrata 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea involucrata 4 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea jacea 2 Ljunghusen, Stora Hammar Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea jacea 3 Ljunghusen, Stora Hammar Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea jacea 4 Ljunghusen, Stora Hammar Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea jacea 6 Imatra Mills FI 120730 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea jacea x nigra 1 Höllviken Sk 130623 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea jacea x nigra 4 Höllviken Sk 130623 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea jacea x nigra 5 Höllviken Sk 130623 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea leucophylla 2 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea leucophylla 3 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea macrocephala 5 Lund Bot Garden 150713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea macrocephala 6 Lund Bot Garden 150713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea melitensis 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea melitensis 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea montana 2 Garda de Sus, Romania 090615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea montana 3 Copenhagen Bot Garden 121120 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea napulifera 1 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea napulifera 2 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea napulifera 3 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea napulifera 4 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea napulifera 5 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea napulifera 6 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nervosa ssp promota 1 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nervosa ssp promota 2 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 1 Kungsladugården, Visby, Gotland 910803 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 11 E of Önneslöv, Bonderup Sk 140820 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea paniculata 2 Comboire FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea paniculata 3 Comboire FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea parilica 1 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea parilica 2 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea parilica 3 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea parilica 4 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea parilica 5 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea parilica 6 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp phrygia 4 Linnansaari National Park FI 120731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp pseudophrygia 4 Böckstein, Austria 080701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp pseudophrygia 5 Lindeskov WSW of Örbaek, Fyn DK 120901 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp pseudophrygia 6 Lindeskov WSW of Örbaek, Fyn DK 120901 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea polyacantha 1 Bolonia, Andalusia, Spain 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea polyacantha 2 Bolonia, Andalusia, Spain 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea polyacantha 3 Bolonia, Andalusia, Spain 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea polyacantha 4 Bolonia, Andalusia, Spain 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea polyacantha 5 Bolonia, Andalusia, Spain 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea polyacantha 6 Bolonia, Andalusia Sp 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pulcherrima 1 (cult) Lund Sk 130623 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pulcherrima 2 (cult) Lund Sk 130623 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 10 Río de la Miel, Los Alcornocales Natural Park, Andalucía Spain 220413 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 11 Río de la Miel, Los Alcornocales Natural Park, Andalucía Spain 220413 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 12 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 13 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 14 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 2 Gibraltar, Great Britain 120321 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 3 Ronda, Andalusia, Spain 120320 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 4 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 5 Gibraltar, Great Britain 120321 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 6 Río de la Miel, Los Alcornocales Natural Park, Andalucía Spain 220413 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 8 Río de la Miel, Los Alcornocales Natural Park, Andalucía Spain 220413 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pumilio 2 3 km W of Paleochora, Crete 800509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea raphanina ssp mixta 1 S of Kratigos, Lesbos, Greece 130512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea raphanina ssp mixta 2 S of Kratigos, Lesbos, Greece 130512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea raphanina ssp mixta 3 S of Kratigos, Lesbos, Greece 130512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea raphanina ssp mixta 4 S of Kratigos, Lesbos, Greece 130512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea raphanina ssp mixta 5 S of Kratigos, Lesbos, Greece 130512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea raphanina ssp raphanina 1 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea raphanina ssp raphanina 2 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea rupestris 2 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea rupestris 3 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea rupestris 4 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea ruthenica 1 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea ruthenica 2 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea ruthenica 3 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea ruthenica 4 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea salonitana 3 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea scabiosa 3 Lund Sk 120702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea scabiosa 4 Nyvik, Stora Hammar, Sk 130808 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea scabiosa 5 Lund Sk 120702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea scabiosa 6 Nyvik, Stora Hammar, Sk 130808 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea simplicicaulis 1 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea simplicicaulis 2 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea solstitialis 3 Sigri, Lesbos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sphaerocephala 3 Cantales de la Araña, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sphaerocephala ssp polyacantha 1 Bolonia, Andalusia, Spain 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sphaerocephala ssp polyacantha 2 Bolonia, Andalusia, Spain 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sphaerocephala ssp polyacantha 3 Bolonia, Andalusia, Spain 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sphaerocephala ssp polyacantha 4 Bolonia, Andalusia, Spain 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sphaerocephala ssp polyacantha 5 Bolonia, Andalusia, Spain 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sphaerocephala ssp polyacantha 6 Bolonia, Andalusia Sp 120324 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea stoebe 1 Lund Bot Garden 130816 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea stoebe 3 Daugavpils stn, Latvia 100712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea stoebe 4 Norra Åsum Sk 110812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea stoebe 5 Norra Åsum Sk 110812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sulphurea 4 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea triumfettii 3 Holomondas, Greece 050528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea triumfettii 4 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea uniflora ssp uniflora 1 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea uniflora ssp uniflora 3 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea uniflora ssp uniflora 4 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea urvillei 1 N of Marathokampos, Samos, Greece 920506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea urvillei 2 SE of Megali Limni, Lesbos, Greece 100515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea 1a Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea 1b Copenhagen Bot Garden 160713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea Subgenus Acrolophus 1 near Lake Mikri Prespa, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea Subgenus Acrolophus 2 near Lake Mikri Prespa, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea adamii 4 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea affinis 1 near Milea, old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea affinis 2 near Milea, old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea argentea Impros, Crete 800515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea calocephala 1 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea calocephala 2 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf affinis 1 Paranesti, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf alba 3 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cf nervosa ssp promota 1 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea clementei 1 Sierra del Co, Villanueva del Cauche, Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea cyanus 1 Dalby Västermark Sk 130618 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea diffusa 2 Tallinn, Estonia 100708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea diluta 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea fischeri 1 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 1 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 4 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea graeca 5 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea jacea 1 Ljunghusen, Stora Hammar Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea jacea x nigra 2 Höllviken Sk 130623 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea jacea x nigra 3 Höllviken Sk 130623 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea lainzii 1 Monardilla, Sierra Bermeja, Andalucía Spain 220416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea lainzii 2 Monardilla, Sierra Bermeja, Andalucía Spain 220416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea leucophylla 1 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea macrocephala 1 Lund Bot Garden 150818 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea macrocephala 2 Lund Bot Garden 150818 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea macrocephala 3 Lund Bot Garden 150818 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea macrocephala 4 Lund Bot Garden 150713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea macrocephala 7 Lund Bot Garden 150713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea melitensis 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea melitensis 4 Miradór de Don Martín, Güímar, Tenerife 160405 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea melitensis 5 Miradór de Don Martín, Güímar, Tenerife 160405 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea melitensis 6 Miradór de Don Martín, Güímar, Tenerife 160405 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea montana 1 Garda de Sus, Romania 090615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea montana 4 Copenhagen Bot Garden 121120 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 10 E of Önneslöv, Bonderup Sk 140820 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 2 N of Fårarp, Högestad, Skåne 930807 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 3 N of Fårarp, Högestad, Skåne 930807 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 4 E of Önneslöv, Bonderup Sk 140820 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 5 E of Önneslöv, Bonderup Sk 140820 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 6 E of Önneslöv, Bonderup Sk 140820 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 7 E of Önneslöv, Bonderup Sk 140820 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 8 E of Önneslöv, Bonderup Sk 140820 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea nigra 9 E of Önneslöv, Bonderup Sk 140820 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea paniculata 1 Comboire FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea parilica 7 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea parilica 8 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp phrygia 1 Linnansaari National Park FI 120731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp phrygia 2 Linnansaari National Park FI 120731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp phrygia 3 Lasila, Estonia 100708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp phrygia 5 Lasila, Estonia 100708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp pseudophrygia 1 Brandstrup, Jylland 090830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp pseudophrygia 2 Bad Gastein, Austria 080704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea phrygia ssp pseudophrygia 3 Bad Gastein, Austria 080704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pterocaula 1 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pterocaula 2 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pulcherrima 3 (cult) Lund Sk 130623 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 7 Río de la Miel, Los Alcornocales Natural Park, Andalucía Spain 220413 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pullata 9 Río de la Miel, Los Alcornocales Natural Park, Andalucía Spain 220413 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea pumilio 1 3 km W of Paleochora, Crete 800509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea rupestris 1 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea salonitana 1 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea salonitana 2 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea salonitana 4 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea salonitana 5 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea salonitana 6 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea salonitana 7 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea scabiosa 1 Kävlinge, Skåne 890702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea scabiosa 2 Nyvik, Stora Hammar, Sk 130808 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea simplicicaulis 3 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea solstitialis 1 Crystal Cove State Park, Newport Beach, CA 130414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea solstitialis 2 Crystal Cove State Park, Newport Beach, CA 130414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sphaerocephala 1 Cantales de la Araña, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sphaerocephala 2 Cantales de la Araña, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sphaerocephala 4 Cantales de la Araña, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea stoebe 2 Lund Bot Garden 130816 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sulphurea 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sulphurea 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea sulphurea 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea triumfettii 1 Holomondas, Greece 050528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea triumfettii 2 Holomondas, Greece 050528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea triumfettii 5 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Centaurea uniflora ssp uniflora 2 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).