Cirsium 1a Marais de Chapareillan FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium 2c Col d'Izoard FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium 2d Col d'Izoard FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium acaule 1 Benestad Sk 120816 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium acaule 2 Benestad Sk 120816 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium acaule 3 Botes springmire, Gotland 070804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium appendiculatum 1 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium appendiculatum 2 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium appendiculatum 5 Rodhopi Mountains, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium arvense 3 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium arvense 4 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium arvense 5 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium arvense 6 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium arvense 7 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium canum 1 W of Bara, Skåne 000804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium canum 4 (cult) Brunnby Sk 120828 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium canum 5 (cult) Brunnby Sk 120828 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium canum 6 (cult) Brunnby Sk 120828 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 10 Burren Nat Park, Clare, Ireland 170528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 11 Burren Nat Park, Clare, Ireland 170528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 12 Burren Nat Park, Clare, Ireland 170528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 13 Burren Nat Park, Clare, Ireland 170528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 3 Burren Nat Park, Clare, Ireland 170528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 4 Burren Nat Park, Clare, Ireland 170528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 5 Burren Nat Park, Clare, Ireland 170530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 6 Burren Nat Park, Clare, Ireland 170528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 7 Burren Nat Park, Clare, Ireland 170530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 8 Burren Nat Park, Clare, Ireland 170528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 9 Burren Nat Park, Clare, Ireland 170528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium eriophorum 1 Mont Aigoual, Cévennes Nat. Park, France 180720 ( - Photo: (© 2014 Jan Thomas Johansson).

Cirsium erisithales 3 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium erisithales 4 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 10 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 13 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 14 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 15 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 16 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 17 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 3 Böckstein, Austria 080701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 4 Tiiriönmäki FI 120802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 5 Tiiriönmäki FI 120802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 8 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 9 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium henryi 2 Shika, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium henryi 3 Shika, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium heterophyllum x palustre 5 Harjula, Vuostimojärvi W of Vuostimo, Lappland, Finland 220728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium japonicum var spinosissimum 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium japonicum var spinosissimum 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium japonicum var spinosissimum 2 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium japonicum var spinosissimum 3 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 1 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 2 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 7 Marais de Montfort (Marais de Crolles) FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 8 Marais de Montfort (Marais de Crolles) FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 10 Vesterroder SE of Stokkemarke, Lolland DK 130811 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 3 Vesterroder SE of Stokkemarke, Lolland DK 130811 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 4 Västra Vemmenhög Sk 130907 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 5 Västra Vemmenhög Sk 130907 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 6 Mullerup S Revninge S Kerteminde, Fyn DK 120901 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 7 Västra Vemmenhög Sk 130907 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 8 Vesterroder SE of Stokkemarke, Lolland DK 130811 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 9 Vesterroder SE of Stokkemarke, Lolland DK 130811 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium palustre 1 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium palustre 2 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium palustre 3 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium rivulare 1 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium rivulare 5 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium rivulare 6 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium shansiense 2 Shangri-la, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium spinosissimum 2 Nassfeld, Austria 080701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium spinosissimum 3 Nassfeld, Austria 080701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium spinosissimum 3 between Col du Lautaret and Col du Galibier FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium spinosissimum 4 between Col du Lautaret and Col du Galibier FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium subcoriaceum 4 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium subcoriaceum 6 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium subcoriaceum 7 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium subcoriaceum 8 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium subcoriaceum 9 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium tuberosum 2 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium tuberosum 3 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium tuberosum 4 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium tuberosum 5 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium tuberosum 6 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium vulgare 4 Ljunghusen, Stora Hammar Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium 1b Marais de Chapareillan FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium 2a Col d'Izoard FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium 2b Col d'Izoard FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium appendiculatum 3 Partheno Dasos, Rodhopi Mountains, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium appendiculatum 4 Partheno Dasos, Rodhopi Mountains, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium arvense 1 Lund Sk 130627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium arvense 2 Lund Sk 130627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium brachycephalum 1 Nagyiváni puszta, Hortobágy, Hungary 240511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium brachycephalum 2 Nagyiváni puszta, Hortobágy, Hungary 240511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium canum 2 W of Bara, Skåne 811006 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium canum 3 W of Bara, Skåne 811006 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 1 Burren Nat Park, Clare, Ireland 170530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 14 Burren Nat Park, Clare, Ireland 170528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium dissectum 2 Burren Nat Park, Clare, Ireland 170530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium erisithales 1 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium erisithales 2 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 1 Böckstein, Austria 080701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 11 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 12 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 2 Törnskogen, Sollentuna, Uppland 040812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium helenioides 6 Tiiriönmäki FI 120802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium henryi 1 Shika, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium heterophyllum x palustre 1 Harjula, Vuostimojärvi W of Vuostimo, Lappland, Finland 220728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium heterophyllum x palustre 2 Harjula, Vuostimojärvi W of Vuostimo, Lappland, Finland 220728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium heterophyllum x palustre 3 Harjula, Vuostimojärvi W of Vuostimo, Lappland, Finland 220728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium heterophyllum x palustre 4 Harjula, Vuostimojärvi W of Vuostimo, Lappland, Finland 220728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium japonicum var spinosissimum 1 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 1 Marais de Montfort (Marais de Crolles) FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 2 Marais de Montfort (Marais de Crolles) FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 3 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 3 Marais de Montfort (Marais de Crolles) FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 4 Marais de Montfort (Marais de Crolles) FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 5 Marais de Montfort (Marais de Crolles) FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium monspessulanum 6 Marais de Montfort (Marais de Crolles) FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 1 Västra Vemmenhög Sk 130907 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 11 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 12 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 13 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 14 Önnarp, Hössna, Västergötland 140726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium oleraceum 2 Mullerup S Revninge S Kerteminde, Fyn DK 120901 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium pannonicum 1 E of Veseli, Slovakia 950630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium pannonicum 2 E of Veseli, Slovakia 950630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium rivulare 2 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium rivulare 3 Lund Bot Garden 080610 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium rivulare 4 Lund Bot Garden 080610 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium shansiense 1 Shangri-la, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium spinosissimum 1 Nassfeld, Austria 080701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium spinosissimum 1 between Col du Lautaret and Col du Galibier FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium spinosissimum 2 between Col du Lautaret and Col du Galibier FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium spinosissimum 5 Col d'Izoard FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium spinosissimum 6 Col d'Izoard FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium subcoriaceum 1 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium subcoriaceum 2 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium subcoriaceum 3 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium subcoriaceum 5 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium tuberosum 1 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium vulgare 1 Ljunghusen, Stora Hammar Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium vulgare 2 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium vulgare 3 Ljunghusen, Stora Hammar Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium vulgare 5 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Cirsium vulgare 6 Skåne-Tranås Sk 070706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).