Knautia arvensis ssp arvensis 2 Toftagården, Tofta, Gotland 070802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia arvensis ssp kitaibelii 2 W of Zilina, Slovakia 950630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia cf integrifolia 1 W of Mandra, Thracia, Greece 060605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia cf integrifolia 2 W of Mandra, Thracia, Greece 060605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia cf integrifolia 4 Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia cf integrifolia 5 Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia cf integrifolia 6 Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia cf integrifolia 7 Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia dipsacifolia 5 Bad Gastein, Austria 080630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia dipsacifolia 6 Bad Gastein, Austria 080704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia dipsacifolia 7 Copenhagen Bot Garden 080513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia dipsacifolia ssp dipsacifolia 10 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia dipsacifolia ssp dipsacifolia 9 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia drymeia ssp intermedia 1 Bergianska 060821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia drymeia ssp intermedia 2 Bergianska 060821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia integrifolia 2 Holomondas, Greece 050528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia integrifolia 3 Kallonis, Lesvos, Greece 130506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia integrifolia 4 NW of Dadia, Thracia, Greece 060606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia integrifolia 5 NW of Mandamados, Lesvos, Greece 100514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia macedonica 1 Lund Bot Garden 160721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia macedonica 2 Lund Bot Garden 160721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia macedonica 3 Lund Bot Garden 160721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia macedonica 4 Lund Bot Garden 160721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia macedonica 5 Lund Bot Garden 160721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia macedonica Lund Bot Garden 070720 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia magnifica 3 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia magnifica 5 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia midzorensis 1 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia midzorensis 2 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia midzorensis 3 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia midzorensis 4 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia midzorensis 5 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 1 Bergianska, Stockholm 010824 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 10 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 11 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 2 W of Mandra, Thracia, Greece 060605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 3 between Mikroderio and Metaxades, Thracia, Greece 060608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 4 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 5 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 6 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 7 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia tatarica 4 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia arvensis ssp arvensis 1 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia arvensis ssp kitaibelii 1 W of Zilina, Slovakia 950630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia cf integrifolia 3 Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia dipsacifolia 1 Bad Gastein, Austria 080630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia dipsacifolia 2 Bad Gastein, Austria 080704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia dipsacifolia 3 Bad Gastein, Austria 080704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia dipsacifolia 4 Copenhagen Bot Garden 080513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia dipsacifolia ssp dipsacifolia 8 between Lans en Vercors and Villard-de-Lans FR 150702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia integrifolia 1 Holomondas, Greece 050528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia magnifica 1 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia magnifica 2 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia magnifica 4 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia magnifica 5 Mt Varnous, N Greece 180630 (2) - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 8 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia orientalis 9 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia tatarica 1 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia tatarica 2 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Knautia tatarica 3 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).