Vaccinium angustifolium 2 Storrs, Connecticut 900508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium calycinum 2 Pihea Trail, Alakui Swamps, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium calycinum 5 Pihea Trail, Alakui Swamps, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium calycinum 6 Pihea Trail, Alakui Swamps, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium calycinum 7 Pihea Trail, Alakui Swamps, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium calycinum 8 Pihea Trail, Alakui Swamps, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium cf corymbosum 1 Copenhagen Bot Garden 080513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium cf corymbosum 2 Copenhagen Bot Garden 080513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium cf emirnense 2 S of Antsirabe, Madagascar 191016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium cf emirnense 3 S of Antsirabe, Madagascar 191016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium cf emirnense 4 S of Antsirabe, Madagascar 191016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium cf emirnense 5 S of Antsirabe, Madagascar 191016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium cf sikkimense Yunnan 1107 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium corymbosum 1 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium corymbosum 2 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium corymbosum 3 Lund Bot Garden 130527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium corymbosum 4 Storrs, Connecticut 900501 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium corymbosum 5 Storrs, Connecticut 900515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium dentatum 2 Puu Kaua, Oahu, Hawaii 840226 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium floribundum 1 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium floribundum 10 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium floribundum 2 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium floribundum 3 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium floribundum 4 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium floribundum 6 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium floribundum 7 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium floribundum 8 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium floribundum 9 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus 5 Höllviken Sk 130805 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus 6 Höllviken Sk 130805 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus 7 Höllviken Sk 130805 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus x vitis-idaea 1 E of Grosshamn, Askum, Bohuslän 990717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus x vitis-idaea 3 Lund Bot Garden 150925 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus x vitis-idaea 4 Lund Bot Garden 150925 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus x vitis-idaea 5 Lund Bot Garden 150925 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus x vitis-idaea 6 Lund Bot Garden 150925 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 11 Kilauea Iki, Big Island, Hawaii 130725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 12 Mauna Loa Trail, Big Island, Hawaii 130725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 13 Mauna Loa Trail, Big Island, Hawaii 130725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 14 Haleakala, Maui, Hawaii 130716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 15 Haleakala, Maui, Hawaii 130716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 3 Kilauea, Big Island, Hawaii 130723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 4 Kilauea, Big Island, Hawaii 130723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 5 Kilauea, Big Island, Hawaii 130723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 8 Kilauea Iki, Big Island, Hawaii 130725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 9 Kilauea Iki, Big Island, Hawaii 130725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium stamineum 1 Macedonia State Park, Connecticut 900606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium stamineum 2 Macedonia State Park, Connecticut 900606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium stapfianum Mt Kinabalu, Sabah, Malaysia 2007 - Photo: Marie Widén (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp microphyllum 1 Mt Hassel, Kangerlussuaq, Greenland 140630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp microphyllum 2 Mt Hassel, Kangerlussuaq, Greenland 140630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp microphyllum 3 Tacan, Kangerlussuaq, Greenland 140629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 10 Brännelycke N of Sölvesborg, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 11 Brännelycke N of Sölvesborg, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 12 Brännelycke N of Sölvesborg, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 2 Ryfors SW of Mullsjö, Västergötland 140722 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 3 Ryfors SW of Mullsjö, Västergötland 140722 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 4 Ryfors SW of Mullsjö, Västergötland 140722 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 5 Brännelycke N of Sölvesborg, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 6 Brännelycke N of Sölvesborg, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 7 Brännelycke N of Sölvesborg, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 8 Brännelycke N of Sölvesborg, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 9 Brännelycke N of Sölvesborg, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp minus 1 Lake Helen, Kangerlussuaq, Greenland 140703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp minus 2 Lake Helen, Kangerlussuaq, Greenland 140703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp minus 3 Lake Helen, Kangerlussuaq, Greenland 140703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp minus 4 Lake Helen, Kangerlussuaq, Greenland 140703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp minus 5 Straumsnesaksla, Båtsfjord, Finnmark, Norway 220726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp minus 6 Straumsnesaksla, Båtsfjord, Finnmark, Norway 220726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp minus 7 Straumsnesaksla, Båtsfjord, Finnmark, Norway 220726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp vitis-idaea 1 E of Mullsjö, Västergötland 140530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp vitis-idaea 2 E of Mullsjö, Västergötland 140530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp vitis-idaea 3 E of Mullsjö, Västergötland 140530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp vitis-idaea 4 E of Mullsjö, Västergötland 140530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp vitis-idaea 5 E of Mullsjö, Västergötland 140530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp vitis-idaea 6 E of Mullsjö, Västergötland 140530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp vitis-idaea 7 E of Mullsjö, Västergötland 140530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp vitis-idaea 8 E of Mullsjö, Västergötland 140530 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium vitis-idaea ssp vitis-idaea 9 Mästocka, Halland 930808 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium 1a (cult) Valencia, CA 130413 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium 1b (cult) Valencia, CA 130413 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium angustifolium 1 Storrs, Connecticut 900508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium calycinum 1 Pihea Trail, Alakui Swamps, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium calycinum 3 Pihea Trail, Alakui Swamps, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium calycinum 4 Pihea Trail, Alakui Swamps, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium cf emirnense 1 S of Antsirabe, Madagascar 191016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium cf emirnense 6 S of Antsirabe, Madagascar 191016 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium dentatum 1 Puu Kaua, Oahu, Hawaii 840226 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium floribundum 5 Cerro de la Muerte, Costa Rica 231211 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus 1 Höllviken Sk 080503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus 2 Höllviken Sk 080503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus 3 Höllviken Sk 080503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus 4 Höllviken Sk 080503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus x vitis-idaea 2 E of Grosshamn, Askum, Bohuslän 990717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium myrtillus x vitis-idaea 7 Lund Bot Garden 150925 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 1 Kilauea, Big Island, Hawaii 130723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 10 Kilauea Iki, Big Island, Hawaii 130725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 2 Kilauea, Big Island, Hawaii 130723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 6 Kilauea, Big Island, Hawaii 130723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium reticulatum 7 Kilauea, Big Island, Hawaii 130723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp microphyllum 4 Tacan, Kangerlussuaq, Greenland 140629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp microphyllum 5 Tacan, Kangerlussuaq, Greenland 140629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Vaccinium uliginosum ssp uliginosum 1 Törnskogen, Sollentuna, Uppland 990608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).