Betonica alopecuros 7 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica alopecuros 8 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 1 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 12 Bergianska 060620 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 4 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 8 Lund Bot Garden 080612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 9 Lund Bot Garden 080612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 13 Bosarp Sk 150721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 14 Bosarp Sk 150721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 15 Bosarp Sk 150721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 16 Bosarp Sk 150721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 8 SE of Lake Ryvalen, Sollentuna, Uppland 840730 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica alopecuros 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica alopecuros 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica alopecuros 3 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica alopecuros 4 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica alopecuros 5 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica alopecuros 6 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica alopecuros 9 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 10 Bergianska 060620 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 11 Bergianska 060620 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 2 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 3 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 5 Lund Bot Garden 080612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 6 Lund Bot Garden 080612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica macrantha 7 Lund Bot Garden 080612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 1 Kungsmarken, Hardeberga Sk 140720 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 10 Bosarp Sk 150721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 11 Bosarp Sk 150721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 12 Bosarp Sk 150721 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 17 Bjergemark E of Rödbyhavn, Lolland DK 160731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 18 Bjergemark E of Rödbyhavn, Lolland DK 160731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 19 Bjergemark E of Rödbyhavn, Lolland DK 160731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 2 Kungsmarken, Hardeberga Sk 140720 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 20 Bjergemark E of Rödbyhavn, Lolland DK 160731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 3 near Lazdukalni, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 4 near Lazdukalni, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 5 near Lazdukalni, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 6 Kungsmarken, Södra Sandby, Skåne 840822 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 7 SE of Lake Ryvalen, Sollentuna, Uppland 840730 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica officinalis 9 SE of Lake Ryvalen, Sollentuna, Uppland 840730 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica pradica 1 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica pradica 2 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica pradica 3 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica scardica 1 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica scardica 2 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica scardica 3 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica scardica 4 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Betonica scardica 5 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).