Euphrasia L. (Orobanchaceae)

Euphrasia arctica ssp minor 3 Råbjerg Mile S of Skagen, Jylland DK 060805 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf pectinata 5 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf pectinata 6 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf pectinata 7 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf pectinata 8 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf pectinata 9 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf regelii 3 NW of Shangrli-la, Yunnan 110708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia dunensis 1 Bulbjerg, Jylland DK 950808 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia dunensis Bulbjerg, Jylland DK 090829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia hyperborea 6 Ansätten, Jämtland 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia hyperborea 7 Ansätten, Jämtland 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 10 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 11 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 8 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 9 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha x stricta var brevipila 1 southern Fanö, Jylland DK 140802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha x stricta var brevipila 2 southern Fanö, Jylland DK 140802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia minima 5 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia minima 6 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 3 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 4 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 8 Södra Paipis, Sibby Fi 120729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 9 Södra Paipis, Sibby Fi 120729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis 3 Bad Gastein, Austria 080630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 1 Täby Prästängar, Täby, Uppland 960731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 10 Sveaborg FI 120804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 11 Sveaborg FI 120804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 12 Sveaborg FI 120804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 13 Sveaborg FI 120804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 9 Sveaborg FI 120804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 10 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 11 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 4 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 5 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 6 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 7 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia pectinata 1 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia pectinata 2 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia regelii 1 between Lijiang and Shangri-la, Yunnan 110708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia regelii 2 between Lijiang and Shangri-la, Yunnan 110708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp salisburgensis 1 Kopparåsen, Torne Lappmark 110727 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp salisburgensis 2 Kopparåsen, Torne Lappmark 110727 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp salisburgensis 3 Kopparåsen, Torne Lappmark 110727 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp salisburgensis 4 Kopparåsen, Torne Lappmark 110727 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp salisburgensis 8 Lullihatjårro, Torne Lappmark 110728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp schoenicola 6 Botes, Lojsta, Gotland 070804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp schoenicola 7 Botes, Lojsta, Gotland - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp schoenicola 8 Botes, Lojsta, Gotland - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp schoenicola 9 Botes, Lojsta, Gotland - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var schoenicola 10 Botes Källmyr, Lojsta, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var schoenicola 17 Botes Källmyr, Lojsta, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var schoenicola 18 Botes Källmyr, Lojsta, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var schoenicola 19 Botes Källmyr, Lojsta, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta 1 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta 2 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta 3 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta 4 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta 5 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 10 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 11 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 4 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 5 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 6 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 7 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 8 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila Gnisvärd, Tofta, Gotland 070802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var gotlandica 1 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var gotlandica 2 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var gotlandica 6 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var gotlandica 7 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var gotlandica 8 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var gotlandica 9 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 13 E of Langmyr, Östergarn, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 5 Kaugatoma-Loo alvar, Saaremaa, Estonia 100717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 6 E of Langmyr, Östergarn, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 7 E of Langmyr, Östergarn, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 8 E of Langmyr, Östergarn, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var suecica 1 Alvena Lindaränge, Vallstena, Gotland 800622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var suecica 3 Anga Prästänge, Gotland 830626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var suecica 4 Anga Prästänge, Gotland 070604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var suecica 5 Anga Prästänge, Gotland 070604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 11 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 12 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 13 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 14 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 15 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 16 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 17 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 2 2 km SW of Landön, Trolle-Ljungby, Skåne 940710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 6 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 1 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 10 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 11 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 4 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 7 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 8 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 9 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii var palustris 1 Ansätten, Jämtland 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii var palustris 2 Ansätten, Jämtand 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii var palustris 3 Ansätten, Jämtand 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii var palustris 4 Ansätten, Jämtand 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii var palustris 5 Ansätten, Jämtand 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).


Euphrasia arctica ssp borealis Trosavik, S of Stavanger, Norway 050805 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia arctica ssp minor 1 Råbjerg Mile S of Skagen, Jylland DK 060805 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia arctica ssp minor 2 Råbjerg Mile S of Skagen, Jylland DK 060805 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia arctica ssp minor 4 Skagen, Jylland DK 090829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia arctica ssp minor 5 Skagen, Jylland DK 090829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia arctica ssp minor 6 Skagen, Jylland DK 090829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia borneensis Mt Kinabalu, Sabah, Malaysia 2007 - Photo: Marie Widén (© 2014 Jan Thomas Johansson).

Euphrasia borneensis Mt Kinabalu, Sabah, Malaysia 2007 Photo: Marie Widén (© 2014 Jan Thomas Johansson).

Euphrasia cf hirtella 1 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf hirtella 2 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf pectinata 1 between Mikroderio and Metaxades, Thracia, Greece 060608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf pectinata 2 between Mikroderio and Metaxades, Thracia, Greece 060608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf pectinata 3 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf pectinata 4 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf regelii 1 NW of Shangrli-la, Yunnan 110708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf regelii 2 NW of Shangrli-la, Yunnan 110708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia cf scottica Kjörsvik, Norway 820805 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia dunensis 2 Bulbjerg, Jylland DK 950808 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia dunensis 3 Bulbjerg, Jylland DK 950808 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia hyperborea 1 Ansätten, Jämtland 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia hyperborea 2 Ansätten, Jämtland 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia hyperborea 3 Ansätten, Jämtland 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia hyperborea 4 Ansätten, Jämtland 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia hyperborea 5 Ansätten, Jämtland 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia hyperborea 8 Ansätten, Jämtland 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 1 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 2 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 3 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 4 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 5 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 6 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia micrantha 7 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia minima 1 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia minima 2 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia minima 3 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia minima 4 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 1 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 10 Södra Paipis, Sibby Fi 120729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 11 Södra Paipis, Sibby Fi 120729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 2 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 5 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 6 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia nemorosa 7 Skanörs Ljung Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis 1 Bad Gastein, Austria 080630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis 2 Bad Gastein, Austria 080630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 2 Täby Prästängar, Täby, Uppland 960731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 3 Täby Prästängar, Täby, Uppland 960731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 4 Viidumäe, Saaremaa, Estonia 050719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 5 Viidumäe, Saaremaa, Estonia 050719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 6 Sveaborg FI 120804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 7 Sveaborg FI 120804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp officinalis 8 Sveaborg FI 120804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 1 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 12 Dagstorp, Skåne 890908 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 2 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 3 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 8 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia officinalis ssp pratensis 9 Skogastorpskärren, Högstena, Västergötland 140724 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia pectinata 3 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp salisburgensis 5 Lullihatjårro, Torne Lappmark 110728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp salisburgensis 6 Lullihatjårro, Torne Lappmark 110728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp salisburgensis 7 Lullihatjårro, Torne Lappmark 110728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp salisburgensis 9 Kopparåsen, Torne Lappmark 110727 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp schoenicola 1 Botes, Lojsta, Gotland 070804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp schoenicola 2 Botes, Lojsta, Gotland 070804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp schoenicola 3 Botes, Lojsta, Gotland 070804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp schoenicola 4 Botes, Lojsta, Gotland 070804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis ssp schoenicola 5 Botes, Lojsta, Gotland - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var salisburgensis 10 La Meije near Col du Lautaret FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var salisburgensis 11 La Meije near Col du Lautaret FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var salisburgensis 12 La Meije near Col du Lautaret FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var salisburgensis 13 La Meije near Col du Lautaret FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var schoenicola 11 Botes Källmyr, Lojsta, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var schoenicola 12 Botes Källmyr, Lojsta, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var schoenicola 13 Botes Källmyr, Lojsta, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var schoenicola 14 Botes Källmyr, Lojsta, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var schoenicola 15 Botes Källmyr, Lojsta, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia salisburgensis var schoenicola 16 Botes Källmyr, Lojsta, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia scottica Trosavik, S of Stavanger, Norway 050805 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 1 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 12 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 13 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 2 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 3 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var brevipila 9 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var gotlandica 3 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var gotlandica 4 Stigmyr, Hangvar, Gotland - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var gotlandica 5 Stigmyr, Hangvar, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 1 W of Stigmyr, Hangvar, Gotland 070803 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 10 E of Langmyr, Östergarn, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 11 E of Langmyr, Östergarn, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 12 E of Langmyr, Östergarn, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 2 W of Stigmyr, Hangvar, Gotland 070803 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 3 Norra Gattet, Fleringe, Gotland 070803 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 4 Kaugatoma-Loo alvar, Saaremaa, Estonia 100717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var stricta 9 E of Langmyr, Östergarn, Gotland 140830 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 1 2 km SW of Landön, Trolle-Ljungby, Skåne 940710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 10 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 7 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 8 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis 9 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis Lasses äng N of Hamra, Sundre, Gotland 800621 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia stricta var tenuis Östergarns Änge, Gotland 830627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 2 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 3 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 5 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii 6 Store Saltsö, Kangerlussuaq, Greenland 140708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Euphrasia wettsteinii var palustris 6 Ansätten, Jämtand 090704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).