Melampyrum arvense 10 S of Rinkaby, Skåne 940710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 15 Everöd Sk 190709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 16 Everöd Sk 190709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 17 Everöd Sk 190709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 5 Gnisvärd, Tofta, Gotland 070802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 6 Gnisvärd, Tofta, Gotland 070802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 7 Transval, Åhus Sk 140601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 8 Transval, Åhus Sk 140601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 9 Transval, Åhus Sk 140601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 4 Tihany Peninsula, Balaton-felvidék, Hungary 24050 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 5 Tihany Peninsula, Balaton-felvidék, Hungary 24050 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 6 Tihany Peninsula, Balaton-felvidék, Hungary 24050 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 7 Tihany Peninsula, Balaton-felvidék, Hungary 24050 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 8 Tihany Peninsula, Balaton-felvidék, Hungary 24050 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 10 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 11 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 12 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 13 Sandhammaren, Löderup Sk 210902 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 14 Sandhammaren, Löderup Sk 210902 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 15 Sandhammaren, Löderup Sk 210902 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 16 Sandhammaren, Löderup Sk 210902 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 3 W of Lenstad, Torslunda, Öland 080614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 4 W of Lenstad, Torslunda, Öland 080614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 5 W of Lenstad, Torslunda, Öland 080614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 6 W of Lenstad, Torslunda, Öland 080614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 7 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum heracleoticum 3 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp bihariense 1 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp bihariense 2 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp nemorosum 1 Fjälkestad, Skåne 820702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp nemorosum 6 Nohlmarken, Skultorp, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp nemorosum 7 Nohlmarken, Skultorp, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp nemorosum 9 Lilla Frescati, Stockholm, Uppland 060913 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp polonicum 2 Torasoo, Rapla, Estonia 100718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp polonicum 3 Torasoo, Rapla, Estonia 100718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp polonicum 4 Torasoo, Rapla, Estonia 100718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp polonicum 5 Torasoo, Rapla, Estonia 100718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 10 Holmudden, Fårö, Gotland 070803 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 4 Pålsjö skog, Helsingborg Sk 140604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 5 Kallviksudden (Kallavi) FI 120728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 6 Kallviksudden (Kallavi) FI 120728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 7 Kallviksudden (Kallavi) FI 120728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 9 Holmudden, Fårö, Gotland 070803 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum sylvaticum 1 Marknadsbacken, Abborrforsen, Kymmene älv FI 120729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum sylvaticum 2 Marknadsbacken, Abborrforsen, Kymmene älv FI 120729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum sylvaticum 3 Marknadsbacken, Abborrforsen, Kymmene älv FI 120729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum sylvaticum 4 Marknadsbacken, Abborrforsen, Kymmene älv FI 120729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum sylvaticum 5 Marknadsbacken, Abborrforsen, Kymmene älv FI 120729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 1 Gnisvärd, Tofta, Gotland 070802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 11 W of Dämbaträsk, Fårö, Gotland 830625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 12 Everöd Sk 190709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 13 Everöd Sk 190709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 14 Everöd Sk 190709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 2 Gnisvärd, Tofta, Gotland 070802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 3 Transval, Åhus Sk 140601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum arvense 4 Transval, Åhus Sk 140601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 1 Tihany Peninsula, Balaton-felvidék, Hungary 240506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 10 Hosszúhetény, Nagy-mezö, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 11 Hosszúhetény, Nagy-mezö, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 2 Tihany Peninsula, Balaton-felvidék, Hungary 24050 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 3 Tihany Peninsula, Balaton-felvidék, Hungary 24050 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum barbatum 9 Hosszúhetény, Nagy-mezö, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 1 W of Lenstad, Torslunda, Öland 080614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 2 W of Lenstad, Torslunda, Öland 080614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 8 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum cristatum 9 Horns Löväng, Persnäs, Öland 150605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum heracleoticum 1 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum heracleoticum 2 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp bihariense 3 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp bihariense 4 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp bihariense 5 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp nemorosum 2 Nohlmarken, Skultorp, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp nemorosum 3 Nohlmarken, Skultorp, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp nemorosum 4 Nohlmarken, Skultorp, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp nemorosum 5 Nohlmarken, Skultorp, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp nemorosum 8 Nohlmarken, Skultorp, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum nemorosum ssp polonicum 1 Torasoo, Rapla, Estonia 100718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 1 Pålsjö skog, Helsingborg Sk 140604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 11 Valön, Bohuslän 900704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 2 Pålsjö skog, Helsingborg Sk 140604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 3 Pålsjö skog, Helsingborg Sk 140604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum pratense 8 Holmudden, Fårö, Gotland 070803 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum sylvaticum 6 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum sylvaticum 7 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Melampyrum sylvaticum 8 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).