Galium (Asperula arcadiensis) 1 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula arcadiensis) 3 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula arcadiensis) 4 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula arcadiensis) 5 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula arcadiensis) 6 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula arcadiensis) 8 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula aristata ssp condensata) 1 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula aristata ssp condensata) 2 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula aristata ssp thessala) 1 Paranesti, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula aristata ssp thessala) 2 Paranesti, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula hexaphylla) 1 Lund Bot Garden 130816 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula hexaphylla) 4 Lund Bot Garden 070625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula hexaphylla) 5 Lund Bot Garden 070625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula hexaphylla) 9 Lund Bot Garden 160615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula orientalis) 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula orientalis) 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula orientalis) 4 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula orientalis) 5 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula orientalis) 6 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula orientalis) 7 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula orientalis) 8 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula scutellaris) 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula scutellaris) 4 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula scutellaris) 5 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula scutellaris) 6 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula scutellaris) 7 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula scutellaris) 8 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula scutellaris) 9 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula suavis) 1 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula suavis) 2 Göteborg Bot Garden 120628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Cruciata glabra) 2 Valea Morii, Romania 090617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Cruciata glabra) 3 Valea Morii, Romania 090617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Cruciata glabra) 4 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 11 south of Segovia SP 140505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 13 NE of Ängstorp, Silvåkra Sk 140906 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 14 NE of Ängstorp, Silvåkra Sk 140906 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 15 NE of Ängstorp, Silvåkra Sk 140906 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 16 NE of Ängstorp, Silvåkra Sk 140906 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 17 NE of Ängstorp, Silvåkra Sk 140906 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 18 NE of Ängstorp, Silvåkra Sk 140906 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 19 NE of Ängstorp, Silvåkra Sk 140906 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 2 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 20 NE of Ängstorp, Silvåkra Sk 140906 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 21 NE of Ängstorp, Silvåkra Sk 140906 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 4 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 5 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 6 Lund Botanical Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 7 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 8 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Valantia muralis) 5 Cantarriján beach near Málaga, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 1a De Hoop, Western Cape, South Africa 100921 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 1b De Hoop, Western Cape, South Africa 100921 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 1c De Hoop, Western Cape, South Africa 100921 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 1d De Hoop, Western Cape, South Africa 100921 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 1e De Hoop, Western Cape, South Africa 100921 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 2a Santa Catalina Island, CA 130407 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 2b Santa Catalina Island, CA 130407 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 2c Santa Catalina Island, CA 130407 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 4a NW of Dadia, Thracia, Greece 060606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 4b NW of Dadia, Thracia, Greece 060606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 5a Santa Margarita River, CA 130415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 5b Santa Margarita River, CA 130415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 5c Santa Margarita River, CA 130415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 6b (cult) Stångby, Skåne 050717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium album 3 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium album 4 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium album 5 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 1 Triumphal Park, CA 130410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 10 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 11 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 12 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 13 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 14 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 15 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 2 Triumphal Park, CA 130410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 3 Triumphal Park, CA 130410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 4 Triumphal Park, CA 130410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 5 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 6 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 9 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium anisophyllon 1 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium anisophyllon 2 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium anisophyllon 3 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium anisophyllon 4 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium aparine 1 Lund Sk 120702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium aparine 2 Lund Sk 120702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium aparine 3 Lund Sk 120702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium aparine 5 Lund Sk 120702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium aparine 6 Höllviken Sk 120718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium aparine 7 Höllviken Sk 120718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium aparine 8 Höllviken Sk 120718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium aparine 9 Höllviken Sk 120718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium arvense 1 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium arvense 2 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium arvense 3 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium arvense 4 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium arvense 5 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium arvense 6 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium arvense 7 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium boreale 5 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium boreale 6 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium boreale 7 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium boreale 8 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf anisophyllon 1 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf anisophyllon 2 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf heldreichii 3 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf heldreichii 4 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf heldreichii 5 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf parisiense 3 Loarre Castle, N Spain 180526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf tomentosum 1 SW of Oudtshoorn, Western Cape, South Africa 100926 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf tomentosum 3 SW of Oudtshoorn, Western Cape, South Africa 100926 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf tomentosum 4 SW of Oudtshoorn, Western Cape, South Africa 100926 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf trichocarpum 3 near Farellones, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf trichocarpum 4 near Farellones, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf trichocarpum 5 near Farellones, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf trichocarpum 6 near Farellones, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf trichocarpum 7 near Farellones, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium circaezans 2 Storrs, Connecticut 900617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium crespianum 1 Cabo Formentor, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium crespianum 2 Cabo Formentor, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium crespianum 5 Cabo Formentor, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium crespianum 6 Cabo Formentor, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium crespianum 8 Cabo Formentor, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 10 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 11 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 13 Valea Morii, Romania 090617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 15 Valea Morii, Romania 090617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 16 Black head, Clare, Ireland 170529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 17 Black head, Clare, Ireland 170529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 18 Black head, Clare, Ireland 170529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 19 Black head, Clare, Ireland 170529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 4 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 5 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 6 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 7 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 8 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 9 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium debile 4 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium debile 5 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium debile 6 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium degenii 6 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium degenii 7 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium divaricatum 1 SSE of Pterounda, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium divaricatum 2 SSE of Pterounda, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hispidum 5 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hispidum 6 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hispidum 7 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hispidum 8 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium lanceolatum 2 Storrs, Connecticut 900617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 10 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 11 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 4 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 5 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 6 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 7 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 8 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium murale 9 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium obliquum 1 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium obliquum 2 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium obliquum 3 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium obliquum 4 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium odoratum 3 Röddinge Sk 080601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 1 Kvinnsgröta, Gräsgård, Öland 150606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 10 W of Möckelmossen, Resmo, Öland 220604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 14 W of Möckelmossen, Resmo, Öland 220604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 15 W of Möckelmossen, Resmo, Öland 220604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 2 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 3 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 5 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 6 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 8 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 9 W of Möckelmossen, Resmo, Öland 220604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium palustre 1 Skanörs Ljung Sk 130608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium palustre 2 Skanörs Ljung Sk 130608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium palustre 3 Skanörs Ljung Sk 130608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium palustre 4 Skanörs Ljung Sk 130608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium palustre 5 Skanörs Ljung Sk 130608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium palustre 6 Lyngsjö Sk 140601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium palustre 7 Lyngsjö Sk 140601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium palustre 8 Lyngsjö Sk 140601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium parisiense 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium parisiense 4 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium parisiense 5 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium parisiense 6 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium parisiense 7 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pedemontanum 1 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pedemontanum 3 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pedemontanum 4 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pedemontanum 6 Tihany Peninsula, Balaton-felvidék, Hungary 240506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pseudoaristatum 1 Monis Gorgianis near old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pseudoaristatum 2 Monis Gorgianis near old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pseudoaristatum 3 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pseudoaristatum 4 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pseudohelveticum 1 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pseudohelveticum 2 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pseudohelveticum 3 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pseudohelveticum 5 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 1 Nassfeld, Austria 080701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 10 between Lans en Vercors and Villard-de-Lans FR 150702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 11 between Lans en Vercors and Villard-de-Lans FR 150702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 12 between Lans en Vercors and Villard-de-Lans FR 150702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 2 Nassfeld, Austria 080701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 3 Nassfeld, Austria 080701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 4 Lund Bot Garden, Skåne 850615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 5 Kulla-Gunnarstorp, Allerum, Skåne 810614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 8 between Lans en Vercors and Villard-de-Lans FR 150702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum coll Zdiar, Slovakia 950629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rivale 10 at Daugava near Aizkraukle, Latvia 060710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rivale 4 at Daugava near Aizkraukle, Latvia 060710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rivale 5 at Daugava near Aizkraukle, Latvia 060710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rivale 6 at Daugava near Aizkraukle, Latvia 060710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rivale 7 at Daugava near Aizkraukle, Latvia 060710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rivale 8 at Daugava near Aizkraukle, Latvia 060710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rivale 9 at Daugava near Aizkraukle, Latvia 060710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 1 Tofta skjutfält, Gotland 980729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 10 Lummelunda, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 11 Lummelunda, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 18 NW of Etebols, Lummelunda, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 19 NW of Etebols, Lummelunda, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 2 Tofta skjutfält, Gotland 980729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 20 NW of Etebols, Lummelunda, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 21 S of Stavklint, Tofta, Gotland 170630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 23 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 24 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 3 E of Halner, Bro, Gotland 830620 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 7 Lummelunda, Gotland 070803 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 9 Lummelunda, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rubioides 2 Körös-Maros Nat Park, Hungary 240510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium saxatile 1 Fågeltofta, Skåne 910621 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium saxatile 2 Mölle, Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium saxatile 3 Mölle, Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium saxatile 4 Mölle, Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium saxatile 5 Mölle, Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium saxatile 6 Mölle, Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium saxatile 7 Mölle, Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 1 near Lazdukalni, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 10 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 11 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 2 near Lazdukalni, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 3 near Lazdukalni, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 4 near Lazdukalni, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 5 near Lazdukalni, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 6 near Lazdukalni, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 7 near Lazdukalni, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 8 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium schultesii 9 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium setaceum 1 SE of Aperi, Karpathos, Greece 040519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium setaceum 2 along the river NE of Gazi Pasa, Turkey 090408 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium setaceum 3 along the river NE of Gazi Pasa, Turkey 090408 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium setaceum 4 along the river NE of Gazi Pasa, Turkey 090408 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sherardiiflorum 1 Katharo, W of Kritsa, Crete 800507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sherardiiflorum 4 NW of Seki, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sherardiiflorum 5 NW of Seki, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium spurium 6 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium spurium 7 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 10 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 11 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 12 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 13 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 14 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 15 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 16 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 17 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 18 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 19 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 20 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 21 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 22 Aran Islands, Clare, Ireland 170527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 23 Aran Islands, Clare, Ireland 170527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 24 Aran Islands, Clare, Ireland 170527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 25 Aran Islands, Clare, Ireland 170527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 26 Aran Islands, Clare, Ireland 170527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 5 Vandplasken, Jylland DK 060806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 9 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium stylosum 1 Lund Bot Garden 090911 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium stylosum 3 Lund Bot Garden 120615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium stylosum 4 Lund Bot Garden 120615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium stylosum 5 Lund Bot Garden 120615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium stylosum 6 Lund Bot Garden 120615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium stylosum 7 Lund Bot Garden 090911 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 1 Vånga, Skåne 930605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 13 Tosthult, Örkened Sk 150716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 14 Tosthult, Örkened Sk 150716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 15 Tosthult, Örkened Sk 150716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 19 Tosthult, Örkened Sk 160617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 20 Tosthult, Örkened Sk 160617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 21 Tosthult, Örkened Sk 170605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 27 Tosthult, Örkened Sk 170605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 28 Tosthult, Örkened Sk 170605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 5 1,4 km E of Gökadal, Blekinge 110814 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 8 1,4 km E of Gökadal, Blekinge 110814 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 1 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 10 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 11 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 12 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 13 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 14 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 15 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 16 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 17 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 2 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 3 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 4 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 6 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 7 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 8 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 9 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium taurinum 2 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium taurinum 3 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium taurinum 4 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium taurinum 5 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium taurinum 8 above Pécs, Mecsek, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium tenuissimum 1 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium tenuissimum 2 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 10 Kvinnsgröta, Gräsgård, Öland 150606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 3 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 4 (cult, orig Västergötland) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 6 Varholmen, Dala, Västergötland 770623 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 9 Kvinnsgröta, Gräsgård, Öland 150606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium tricornutum 1 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium tricornutum 2 Lastos, Karpathos, Greece 040522 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium trifidum 1 Raubergstulen, Norway 820731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium trifidum 2 Raubergstulen, Norway 820731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium trifidum 3 Mellomsjön, Västergötland 870705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triflorum 1 ENE of Peura, Lappland, Finland 220728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triflorum 2 ENE of Peura, Lappland, Finland 220728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triflorum 3 ENE of Peura, Lappland, Finland 220728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triflorum 4 ENE of Peura, Lappland, Finland 220728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium uliginosum 1 Sjöängen, Slöta, Västergötland 140723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium uliginosum 2 Sjöängen, Slöta, Västergötland 140723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium uliginosum 3 Sjöängen, Slöta, Västergötland 140723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium uliginosum 4 Sjöängen, Slöta, Västergötland 140723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium uliginosum 6 Sjöängen, Slöta, Västergötland 140723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium uliginosum 7 Sjöängen, Slöta, Västergötland 140723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium vaillantii Kristianstad, Skåne 940710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valantioides 1 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valantioides 4 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valantioides 5 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valantioides 6 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valantioides 7 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valantioides 8 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 10 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 11 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 12 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 13 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 14 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 6 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 7 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 9 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verrucosum 10 Pantallica area, Sicily 190417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verrucosum 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verrucosum 4 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verrucosum 7 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verrucosum 8 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verrucosum 9 Pantallica area, Sicily 190417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 1 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 10 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 11 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 2 Höllviken Sk 130805 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 4 Höllviken Sk 120717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 6 Höllviken Sk 120717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 7 Höllviken Sk 120717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 8 Höllviken Sk 120717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 9 Skanörs Ljung Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium vexans 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium vexans 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium vexans 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium viridiflorum 2 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium viridiflorum 3 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium viridiflorum 4 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula arcadiensis) 2 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula arcadiensis) 7 Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula aristata ssp thessala) 3 Paranesti, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula aristata ssp thessala) 4 Paranesti, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula hexaphylla) 2 Lund Bot Garden 070625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula hexaphylla) 3 Lund Bot Garden 130610 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula hexaphylla) 6 Lund Bot Garden 160615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula hexaphylla) 7 Lund Bot Garden 160615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula hexaphylla) 8 Lund Bot Garden 160615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula orientalis) 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula rigida) S of Kritsa, Crete 800507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula scutellaris) 1 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Asperula scutellaris) 2 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 10 near Milea, old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 11 near Milea, old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 3 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 4 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 5 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 6 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 7 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 8 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella angustifolia) 9 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella latifolia) 1 SE of Megali Limni, Lesvos, Greece 100515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella latifolia) 2 SE of Megali Limni, Lesvos, Greece 100515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella latifolia) 3 SE of Megali Limni, Lesvos, Greece 100515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella latifolia) 4 SE of Megali Limni, Lesvos, Greece 100515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella latifolia) 5 SE of Megali Limni, Lesvos, Greece 100515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Crucianella latifolia) 6 SE of Megali Limni, Lesvos, Greece 100515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Cruciata glabra) 1 Zdiar, Slovakia 950629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 1 Lund Bot Garden 130825 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 10 south of Segovia SP 140505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 12 Lund Bot Garden 130825 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 3 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Sherardia arvensis) 9 south of Segovia SP 140505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Valantia muralis) 1 Cantarriján beach near Málaga, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Valantia muralis) 2 Cantarriján beach near Málaga, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Valantia muralis) 3 Cantarriján beach near Málaga, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium (Valantia muralis) 4 Cantarriján beach near Málaga, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 3a Suatu, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 3b Suatu, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 3c Suatu, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium 6a (cult) Stångby, Skåne 050717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium aff mollugo N of Gerakini, Greece 050525 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium album 1 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium album 2 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium album x verum 1 Bräcke, Bohuslän 820726 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium album x verum 2 Otta, Norway 820729 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 7 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium angustifolium 8 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium anisophyllon 1 Chamrousse FR 150703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium anisophyllon 2 Chamrousse FR 150703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium anisophyllon 3 Chamrousse FR 150703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium anisophyllon 4 Chamrousse FR 150703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium aparine 4 Lund Sk 120702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium boreale 1 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium boreale 2 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium boreale 3 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium boreale 4 E of Lund Sk 130614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf heldreichii 1 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf heldreichii 2 Mt Falakron, N Greece 180703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf parisiense 1 Loarre Castle, N Spain 180526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf parisiense 2 Loarre Castle, N Spain 180526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf parisiense 4 Loarre Castle, N Spain 180526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf tomentosum 2 SW of Oudtshoorn, Western Cape, South Africa 100926 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf trichocarpum 1 near Farellones, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cf trichocarpum 2 near Farellones, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium circaezans 1 Storrs, Connecticut 900617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium crespianum 3 Cabo Formentor, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium crespianum 4 Cabo Formentor, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium crespianum 7 Cabo Formentor, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cruciata 1 Ambelos Padroso, Samos, Greece 920508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cruciata 2 Gärdet, Stockholm 040618 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cruciata 3 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cruciata 4 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cruciata 5 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cruciata 6 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cruciata 7 Tihany Peninsula, Balaton-felvidék, Hungary 240506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cruciata 8 Tihany Peninsula, Balaton-felvidék, Hungary 240506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cruciata 9 Tihany Peninsula, Balaton-felvidék, Hungary 240506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 1 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 12 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 14 Valea Morii, Romania 090617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 2 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium cynanchicum 3 Visby, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium debile 1 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium debile 2 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium debile 3 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium degenii 1 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium degenii 2 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium degenii 3 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium degenii 4 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium degenii 5 Copenhagen Bot Garden 130611 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium divaricatum 3 SSE of Pterounda, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium divaricatum 4 SSE of Pterounda, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium divaricatum 5 SSE of Pterounda, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium divaricatum 6 SSE of Pterounda, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hispidum 1 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hispidum 2 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hispidum 3 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hispidum 4 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hispidum 9 Alanya, Turkey 090411 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hoffmeisteri 1 Napa Hai, Yunnan 110709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium hoffmeisteri 2 Napa Hai, Yunnan 110709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium lanceolatum 1 Storrs, Connecticut 900617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium obliquum 5 Moneval NE of Grenoble FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium obliquum 6 Moneval NE of Grenoble FR 150705 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium odoratum 1 Röddinge Sk 080601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium odoratum 2 Röddinge Sk 080601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 11 W of Möckelmossen, Resmo, Öland 220604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 12 W of Möckelmossen, Resmo, Öland 220604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 13 W of Möckelmossen, Resmo, Öland 220604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 4 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium oelandicum 7 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium paradoxum Yunnan 1107 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium parisiense 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium parisiense 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pedemontanum 1 Tihany Peninsula, Balaton-felvidék, Hungary 240506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pedemontanum 2 Tihany Peninsula, Balaton-felvidék, Hungary 240506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pedemontanum 2 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pedemontanum 3 Tihany Peninsula, Balaton-felvidék, Hungary 240506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pedemontanum 4 Tihany Peninsula, Balaton-felvidék, Hungary 240506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pedemontanum 5 Tihany Peninsula, Balaton-felvidék, Hungary 240506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pseudohelveticum 4 Cote 2000 between Moucherolle and Moucherotte FR 150704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 6 Kulla-Gunnarstorp, Allerum, Skåne 810614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium pumilum 9 between Lans en Vercors and Villard-de-Lans FR 150702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rivale 1 at Daugava between Slutiski and Verveli, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rivale 2 at Daugava between Slutiski and Verveli, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rivale 3 at Daugava between Slutiski and Verveli, Latvia 100711 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 12 Lummelunda, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 13 NW of Etebols, Lummelunda, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 14 NW of Etebols, Lummelunda, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 15 NW of Etebols, Lummelunda, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 16 NW of Etebols, Lummelunda, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 17 NW of Etebols, Lummelunda, Gotland 170629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 22 S of Stavklint, Tofta, Gotland 170630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 4 E of Halner, Bro, Gotland 830620 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 5 Lummelunda, Gotland 070803 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 6 Lummelunda, Gotland 070803 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rotundifolium 8 Lummelunda, Gotland 140829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium rubioides 1 Körös-Maros Nat Park, Hungary 240510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sherardiiflorum 2 Katharo, W of Kritsa, Crete 800507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sherardiiflorum 3 NW of Seki, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium spurium 1 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium spurium 2 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium spurium 3 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium spurium 4 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium spurium 5 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 1 Valbjerg Sande, Jylland DK 060806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 2 Valbjerg Sande, Jylland DK 060806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 3 Valbjerg Sande, Jylland DK 060806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 4 Vandplasken, Jylland DK 060806 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 6 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 7 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium sterneri 8 SW of Lögstrup, Jylland DK 160614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium stylosum 2 Lund Bot Garden 120615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 10 Tosthult, Örkened Sk 150716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 11 Tosthult, Örkened Sk 150716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 12 Tosthult, Örkened Sk 150716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 16 Tosthult, Örkened Sk 150716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 17 Tosthult, Örkened Sk 150716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 2 1,4 km E of Gökadal, Blekinge 110814 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 22 Tosthult, Örkened Sk 170605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 23 Tosthult, Örkened Sk 170605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 24 Tosthult, Örkened Sk 170605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 25 Tosthult, Örkened Sk 170605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 26 Tosthult, Örkened Sk 170605 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 3 1,4 km E of Gökadal, Blekinge 110814 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 4 1,4 km E of Gökadal, Blekinge 110814 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 6 1,4 km E of Gökadal, Blekinge 110814 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 7 1,4 km E of Gökadal, Blekinge 110814 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var suecicum 9 Tosthult, Örkened Sk 150716 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium suecicum var vestrogothicum 5 Sparresäter, Lerdala, Västergötland 140529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium taurinum 1 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium taurinum 6 above Pécs, Mecsek, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium taurinum 7 above Pécs, Mecsek, Hungary 240508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium tenuissimum 3 near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 1 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 11 Svartdal, Gammelgarn, Gotland 170628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 12 Svartdal, Gammelgarn, Gotland 170628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 2 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 5 (cult, orig Västergötland) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triandrum 7 Lärbro, Gotland 940624 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium tricornutum 1 Lastos, Karpathos, Greece 040522 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium triflorum 5 ENE of Peura, Lappland, Finland 220728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium uliginosum 5 Sjöängen, Slöta, Västergötland 140723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valantioides 2 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valantioides 3 Copenhagen Bot Garden 130604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 1 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 2 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 3 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 4 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 5 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium valdepilosum 8 Mönsted Kalkbrud W of Viborg, Jylland DK 130629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verrucosum 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verrucosum 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verrucosum 5 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verrucosum 6 Copenhagen Bot Garden 150812 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 3 Höllviken Sk 120717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium verum 5 Höllviken Sk 120717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium vexans 4 Spili, Crete 110329 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium vexans 5 Spili, Crete 110329 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium vexans 6 Spili, Crete 110329 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Galium viridiflorum 1 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).