Anacamptis laxiflora 10 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu 3 Pantallica area, Sicily 190416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 11 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 17 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 7 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 8 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 9 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 1 NE of Spili, Crete 110327 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 12 Pantallica area, Sicily 190416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 17 Sardinia, Italy 230421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 2 NE of Spili, Crete 110327 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 3 NE of Spili, Crete 110327 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 4 SE of Mijas, Andalusia SP 120327 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 5 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 6 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 7 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 12 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 13 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 14 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 15 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 16 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis sancta 4 NE of Skalochori, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis sancta 5 NE of Skalochori, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis boryi 1 Dept of Botany, Stockholm University 020418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis boryi 2 Dept of Botany, Stockholm University 020418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis collina 1 Spili, Crete 110329 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis collina 2 Spili, Crete 110329 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis collina 3 Spili, Crete 110329 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis collina 4 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis collina 5 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis collina 6 Soulia, Crete 110326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp coriophora 1 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp coriophora 2 Botiza, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp coriophora 3 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp coriophora 4 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp fragrans 1 Lastos, Greece 040519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp fragrans 2 Lastos, Greece 040519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp fragrans 3 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp fragrans 4 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp fragrans 5 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp fragrans 6 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp fragrans 7 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis coriophora ssp fragrans 8 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis laxiflora 1 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis laxiflora 11 the Katara Pass, N Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis laxiflora 2 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis laxiflora 3 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis laxiflora 4 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis laxiflora 5 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis laxiflora 6 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis laxiflora 7 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis laxiflora 8 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis laxiflora 9 between Ntipi and Larisos, Lesvos, Greece 100513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu 1 Pantallica area, Sicily 190416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu 2 Pantallica area, Sicily 190416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu 4 Pantallica area, Sicily 190417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu 5 Sicily 190420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu 6 Sicily 190420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu x Neotinea lactea 1 Ficazza area, Sicily 190421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu x Neotinea lactea 2 Ficazza area, Sicily 190421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu x Neotinea lactea 3 Ficazza area, Sicily 190421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu x Neotinea lactea 4 Ficazza area, Sicily 190421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu x Neotinea lactea 5 Ficazza area, Sicily 190421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu x Neotinea lactea 6 Ficazza area, Sicily 190421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis longicornu x Neotinea lactea 7 Ficazza area, Sicily 190421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 1 Kungsmarken, Södra Sandby Sk 140516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 10 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 12 Enerum, Böda, Öland 770607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 13 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 14 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 15 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 16 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 18 S of Sibbaboda, Torhamn, Blekinge 160601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 19 S of Sibbaboda, Torhamn, Blekinge 160601 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 2 Persnäs, Öland 120602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 3 Måryd, Södra Sandby, Skåne 810602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 4 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 5 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio 6 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio ssp caucasica 1 Eagle Gorge Nat. Park, Georgia 170425 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis morio ssp caucasica 2 Eagle Gorge Nat. Park, Georgia 170425 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 1 Hoburgsmyr, Gotland 040704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 1 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 10 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 11 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 12 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 13 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 14 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 15 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 16 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 17 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 18 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 19 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 2 Hoburgsmyr, Gotland 040704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 20 Krakvät, Hablingbo, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 20 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 21 Krakvät, Hablingbo, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 22 Krakvät, Hablingbo, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 23 Krakvät, Hablingbo, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 24 Krakvät, Hablingbo, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 3 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 4 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 5 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 6 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 7 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 8 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris 9 Mästermyr, Mästerby, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris ssp elegans 1 Nagyiváni puszta, Hortobágy, Hungary 240511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris ssp elegans 2 Nagyiváni puszta, Hortobágy, Hungary 240511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis palustris ssp elegans 4 Nagyiváni puszta, Hortobágy, Hungary 240511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 10 E of Dedoplistskaro, Georgia 170425 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 11 Pantallica area, Sicily 190416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 13 Sardinia, Italy 230421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 14 Sardinia, Italy 230421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 15 Sardinia, Italy 230421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 16 Sardinia, Italy 230421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 8 E of Dedoplistskaro, Georgia 170425 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis papilionacea 9 E of Dedoplistskaro, Georgia 170425 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 1 Enerum, Böda, Öland 120703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 10 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 11 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 17 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 18 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 19 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 2 Enerum, Böda, Öland 120703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 20 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 21 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 3 Enerum, Böda, Öland 120703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 4 Enerum, Böda Öland 120703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 5 E of Veseli, Slovakia 950630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 6 Samos City, Samos, Greece 920510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 7 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 8 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis pyramidalis 9 Gnisvärd, Tofta, Gotland 170627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis sancta 1 NE of Skalochori, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis sancta 2 NE of Skalochori, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Anacamptis sancta 3 NE of Skalochori, Lesvos, Greece 100510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).