Matthiola 1a Anden Verde, Gran Canaria 080410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola 1b Anden Verde, Gran Canaria 080410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola arabica 3 E of Dedoplistskaro, Georgia 170425 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola arabica 4 E of Dedoplistskaro, Georgia 170425 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola arabica 5 E of Dedoplistskaro, Georgia 170425 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola cf incana 1 Kampos, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola cf incana 2 Kampos, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola cf incana 3 Kampos, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola cf incana 4 Kampos, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola cf incana 5 Kampos, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola cf incana 6 Kampos, Lesvos, Greece 100512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola cf tricuspidata 3 Skala Eresou, Lesvos, Greece 130510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola cf tricuspidata 4 Skala Eresou, Lesvos, Greece 130510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola fruticulosa 10 Sierra de Mijas, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola fruticulosa 5 Sierra de Mijas, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola fruticulosa 6 Sierra de Mijas, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola fruticulosa 8 Sierra de Mijas, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola fruticulosa 9 Sierra de Mijas, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 1 S of Agadir, Morocco 140220 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 10 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 2 S of Agadir, Morocco 140220 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 3 Mirador de San Antón, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 4 Mirador de San Antón, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 9 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola sinuata Akr. Kastellou, Karpathos, Greece 040521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola tricuspidata 1 S of Pefkochori, Greece 050529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola tricuspidata 2 S of Pefkochori, Greece 050529 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola tricuspidata 3 near Sinis, Cabras, Sardinia, Italy 230417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola tricuspidata 4 near Sinis, Cabras, Sardinia, Italy 230417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola 2a Guaza, La Gomera 060412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola 2b Guaza, La Gomera 060412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola arabica 1 E of Dedoplistskaro, Georgia 170425 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola arabica 2 E of Dedoplistskaro, Georgia 170425 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola cf tricuspidata 1 Skala Eresou, Lesvos, Greece 130510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola cf tricuspidata 2 Skala Eresou, Lesvos, Greece 130510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola fruticulosa 1 Sierra de Mijas, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola fruticulosa 2 Sierra de Mijas, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola fruticulosa 3 Sierra de Mijas, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola fruticulosa 4 Sierra de Mijas, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola fruticulosa 7 Sierra de Mijas, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 11 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 12 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 5 Mirador de San Antón, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 6 Mirador de San Antón, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola parviflora 7 Mirador de San Antón, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola sinuata Arkasa, Karpathos, Greece 040520 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Matthiola tricuspidata 5 near Sinis, Cabras, Sardinia, Italy 230417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).