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Viola L. (Violaceae)

Viola 1a Trigrad gorge-Jagodinska, Teshel, Bulgaria 120508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola 1b Trigrad gorge-Jagodinska, Teshel, Bulgaria 120508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola 1f Trigrad gorge-Jagodinska, Teshel, Bulgaria 120508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola aetolica 1 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola aetolica 2 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola aetolica 3 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola aetolica 6 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba 1 Borgholm, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba 2 Borgholm, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba 4 Borgholm, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba 5 Borgholm, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba 6 Borgholm, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba x hirta 1 Solliden, Räpplinge, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 1 Cantales de la Araña, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 10 Cantales de la Araña, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 11 Cantales de la Araña, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 12 Cantales de la Araña, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 2 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 3 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 4 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 5 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 6 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 7 Barranco de la Coladilla, Andalucía Spain 220410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 8 Cantales de la Araña, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arborescens 9 Cantales de la Araña, Andalucía Spain 220412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arvensis 5 Videholm, Stora Hammar Sk 130608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola banksii 1 Lund Bot Garden 120927 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola banksii 3 Lund Bot Garden 120927 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola banksii 4 Lund Bot Garden 120927 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola banksii 5 Lund Bot Garden 120927 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola biflora 1 Söndre Knutshö, Dovre, Norway 820802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola biflora 4 Schlossalm, Austria 080703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola calcarata 2 between Col du Lautaret and Col du Galibier FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola calcarata 3 between Col du Lautaret and Col du Galibier FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola calcarata 4 between Col du Lautaret and Col du Galibier FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola calcarata 5 Chamrousse FR 150703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola calcarata 8 Chamrousse FR 150703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cameleo 1 Shika, Yunnan 110712 - Photo: Unknown (© 2014 Jan Thomas Johansson).

Viola cameleo 2 Shika, Yunnan 110712 - Photo: Unknown (© 2014 Jan Thomas Johansson).

Viola canadensis 1 (cult) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canadensis 2 (cult) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canadensis 3 (cult) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canadensis 4 (cult) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canina 1 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canina 2 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canina 3 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canina 7 Kämpinge, Räng Sk 130518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canina x riviniana Lillhällom W of Sundsvall, Medelpad 870606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf hirta 1 near Lake Kartsakhi, Georgia 170426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf hirta 2 near Lake Kartsakhi, Georgia 170426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf hirta 3 near Lake Kartsakhi, Georgia 170426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf hirta 4 near Lake Kartsakhi, Georgia 170426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf hirta 5 near Lake Kartsakhi, Georgia 170426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf hirta 6 near Lake Kartsakhi, Georgia 170426 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf riviniana 1 Parque Natural Sierra de Gredos, west of Arenas de San Pedro SP 140506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf riviniana 2 Parque Natural Sierra de Gredos, west of Arenas de San Pedro SP 140506 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf rockiana Yunnan 1107 - Photo: Unknown (© 2014 Jan Thomas Johansson).

Viola cf subandina 1 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 10 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 11 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 13 Res. Nac. Rio de los Cipreses E of Rancagua, Chile 171111 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 2 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 3 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 4 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 5 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 6 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 7 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 8 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 9 Res. Nac. Yerba Loca E of Santiago, Chile 171109 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf velutina 1 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf velutina 2 Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola chamissoniana 2 Iliau Loop Trail, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola chamissoniana 3 Iliau Loop Trail, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola chamissoniana 4 Iliau Loop Trail, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola chamissoniana 5 Iliau Loop Trail, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cheiranthifolia 1 Teide, Tenerife 060414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cheiranthifolia 2 Teide, Tenerife 060414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cheiranthifolia 3 Teide, Tenerife 060414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cheiranthifolia 4 Teide, Tenerife 060414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cheiranthifolia 5 Teide, Tenerife 060414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola collina Alnön, Medelpad 8706067 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola conspersa 1 Storrs, Connecticut 900428 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola conspersa 2 Storrs, Connecticut 900504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola conspersa 3 Storrs, Connecticut 900504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cucullata 3 Storrs, Connecticut 900508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cunninghamii 1 Taranaki (Mt Egmont) NZ 151110 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cunninghamii 2 Taranaki (Mt Egmont) NZ 151110 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cunninghamii 3 Taranaki (Mt Egmont) NZ 151110 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cunninghamii 4 Taranaki (Mt Egmont) NZ 151110 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cunninghamii 5 Taranaki (Mt Egmont) NZ 151110 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola declinata 1 Rodnea, Romania 090612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola doerfleri 3 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola doerfleri 6 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola doerfleri 7 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola doerfleri 8 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola doerfleri 9 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 10 Arontorp, Torslunda, Öland 150607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 13 Karum, Högsrum, Öland 150607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 14 Karum, Högsrum, Öland 150607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 4 Tveta, Torslunda, Öland 010609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 5 Tveta, Torslunda, Öland 010609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 8 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 9 Arontorp, Torslunda, Öland 150607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola epipsila 2 Tjietnjalisjaure, Torne Lappmark 110728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola epipsila 3 Tjietnjalisjaure, Torne Lappmark 110728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola epipsila 4 Tjietnjalisjaure, Torne Lappmark 110728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola epipsila 5 Tjietnjalisjaure, Torne Lappmark 110728 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola eximia 6 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola eximia 7 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola filicaulis 2 Pureora Forest Reserve NZ 151118 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola fimbriatula 1 Storrs, Connecticut 900428 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola fimbriatula 2 Storrs, Connecticut 900515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola fragrans Omalos, Crete 800514 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola ganiatsasii 1 Partheno Dasos, Rhodopi Mountains, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola ganiatsasii 3 Partheno Dasos, Rhodopi Mountains, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola hirta 1 Stenby, Benestad Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola hirta 2 Stenby, Benestad Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola hirta 3 Stenby, Benestad Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola hirta 4 Stenby, Benestad Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola hirta 5 Stenby, Benestad Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola jooi 1 (cult) Brunnby Sk 080815 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola jooi 3 (cult) Brunnby Sk 080815 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola jooi 4 (cult) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola jooi 5 (cult) Brunnby Sk 080815 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola jordanii 1 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola jordanii 3 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola kitaibeliana 1 above Seki, near Syedra, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola kitaibeliana 2 above Seki, near Syedra, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola kitaibeliana 4 above Seki, near Syedra, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola labradorica 1 (cult) Lund Sk 160507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola labradorica 2 (cult) Lund Sk 160507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola labradorica 3 (cult) Lund Sk 160507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola labradorica 4 (cult) Lund Sk 160507 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola lanaiensis 1 Olinda Endangered Plant Facility, East Maui, Hawaii 130718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola lanaiensis 2 Olinda Endangered Plant Facility, East Maui, Hawaii 130718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola lanaiensis 3 Olinda Endangered Plant Facility, East Maui, Hawaii 130718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola lanaiensis 5 Olinda Endangered Plant Facility, East Maui, Hawaii 130718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola lanaiensis 8 Olinda Endangered Plant Facility, East Maui, Hawaii 130718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 1 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 11 Arontorp, Torslunda, Öland 150607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 12 Arontorp, Torslunda, Öland 150607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 13 Arontorp, Torslunda, Öland 150607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 4 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 5 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 6 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 7 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 8 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 9 Röddinge Sk 140501 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola odorata 1 Månstorp church Sk 140406 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola odorata 10 Lund Sk 120405 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola odorata 2 Lund Sk 120405 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola odorata 3 Lund Sk 120416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola odorata 4 Lund Sk 140323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola odorata 5 Lund Sk 140323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola odorata 7 Lund Sk 140323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola odorata 8 Lund Sk 120405 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola odorata 9 Lund Sk 140323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmata 1 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmata 2 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmata 3 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmata 4 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmata 5 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmata 6 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 3 below the observatory, La Palma 070417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 4 below the observatory, La Palma 070417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 6 below the observatory, La Palma 070417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 1 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 10 Rövarekulan, Gudmuntorp Sk 210518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 11 Rövarekulan, Gudmuntorp Sk 210518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 12 Rövarekulan, Gudmuntorp Sk 210518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 2 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 4 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 5 Rövarekulan, Gudmuntorp Sk 210518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 7 Rövarekulan, Gudmuntorp Sk 210518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 8 Rövarekulan, Gudmuntorp Sk 210518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 9 Rövarekulan, Gudmuntorp Sk 210518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pedatifida 1 Lund Bot Garden 100527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pedatifida 2 Lund Bot Garden 100527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pedatifida 3 Lund Bot Garden 100527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pedunculata 3 Triumphal Park, CA 130410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pedunculata 4 Triumphal Park, CA 130410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pedunculata 5 Triumphal Park, CA 130410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pubescens 3 Storrs, Connecticut 900511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 1 E of Vickleby, Öland 750603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 11 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 17 Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 2 E of Vickleby, Öland 750603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 3 Eskilstorps Ängar, Skåne 850602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 4 Eskilstorps Ängar, Skåne 850602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 7 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 8 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 9 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 1 Lund Bot Garden 080422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 10 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 11 Örup Sk 150418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 12 Örup Sk 150418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 13 Örup Sk 150418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 14 Örup Sk 150418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 15 Örup Sk 150418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 16 Örup Sk 150418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 2 Lund Bot Garden 080422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 3 Lund Bot Garden 080422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 5 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 6 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 7 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 9 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana x riviniana Lund Bot Garden 100519 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp anagae 10 Chinobre, Tenerife 090517 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp anagae 2 Chinobre, Tenerife 090517 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp anagae 3 Chinobre, Tenerife 090517 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp anagae 4 Chinobre, Tenerife 090517 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp anagae 5 Chinobre, Tenerife 090517 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp anagae 7 Chinobre, Tenerife 090517 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp anagae 8 Chinobre, Tenerife 090517 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp anagae 9 Chinobre, Tenerife 090517 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp riviniana 1 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp riviniana 3 Tofta, Gotland 110508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp riviniana 4 Tofta, Gotland 110508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp riviniana 5 Tofta, Gotland 110508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp riviniana 6 Glengariff Woods, West Cork, Ireland 170523 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola rupestris 1 S of Vitemölla, Vitaby Sk 140419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola rupestris 10 S of Vitemölla, Vitaby Sk 140419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola rupestris 2 S of Vitemölla, Vitaby Sk 140419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola rupestris 4 S of Vitemölla, Vitaby Sk 140419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola rupestris 7 S of Vitemölla, Vitaby Sk 140419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola rupestris 8 S of Vitemölla, Vitaby Sk 140419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola rupestris 9 S of Vitemölla, Vitaby Sk 140419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola scorpiuroides 1 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola scorpiuroides 3 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola scorpiuroides 4 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola selkirkii 2 Halsviksravinen, Ångermanland 870607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola sororia 1 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola sororia 10 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola sororia 2 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola sororia 3 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola sororia 5 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola sororia 6 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola sororia 7 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola sororia 8 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola sororia 9 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 1 Lenstad, Torslunda, Öland 120603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 11 South of Gustav Adolf Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 5 South of Gustav Adolf Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 6 South of Gustav Adolf Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 7 South of Gustav Adolf Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 9 South of Gustav Adolf Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola striata 3 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola striata 4 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola striata 5 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola striata 6 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola striata 8 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola trichopetala 1 Dali, Yunnan 110706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp curtisii 1 Kämpinge, Räng Sk 080503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp curtisii 2 Kämpinge, Räng Sk 080503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp curtisii 3 Kämpinge, Räng Sk 080503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp tricolor 1 Häverö Prästäng, Uppland 980702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp tricolor 2 Häverö Prästäng, Uppland 980702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola uliginosa 3 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola uliginosa 5 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola uliginosa 6 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola uliginosa 8 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola 3 Mt Kaindi, Papua New Guinea 831028 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola adunca 1 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola adunca 2 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola adunca 3 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola adunca 4 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola aetolica 4 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola aetolica 5 above Metsovon, Greece 180627 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba 3 Borgholm, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba 7 Borgholm, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba x hirta 2 Solliden, Räpplinge, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba x odorata 1 Borgholm, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola alba x odorata 2 Borgholm, Öland 890508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arvensis 1 Skanör Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arvensis 2 Videholm, Stora Hammar Sk 130608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arvensis 3 Skanör Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arvensis 4 Skanör Sk 130622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arvensis 6 Nyagrop, Maglehem Sk 140501 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arvensis 7 Nyagrop, Maglehem Sk 140501 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola arvensis 8 Nyagrop, Maglehem Sk 140501 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola banksii 2 Lund Bot Garden 120927 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola biflora 2 Schlossalm, Austria 080703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola biflora 3 Schlossalm, Austria 080703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola blanda Storrs, Connecticut 900512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola calcarata 1 between Col du Lautaret and Col du Galibier FR 150708 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola calcarata 6 Chamrousse FR 150703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola calcarata 7 Chamrousse FR 150703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canadensis 5 Bergianska, Stockholm 050531 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canadensis 6 Bergianska, Stockholm 050531 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canina 4 Kungsmarken, Södra Sandby Sk 110510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canina 5 Kämpinge, Räng Sk 130518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canina 6 Kämpinge, Räng Sk 130518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canina 8 Hammar, Valleberga Sk 120526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola canina 9 Hammar, Valleberga Sk 120526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf subandina 12 Res. Nac. Rio de los Cipreses E of Rancagua, Chile 171111 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf urophylla 1 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cf urophylla 2 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola chamissoniana 1 Iliau Loop Trail, Kauai, Hawaii 130713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola corsica 1 Limbara, Sardinia, Italy 230421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola corsica 2 Limbara, Sardinia, Italy 230421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola corsica 3 Limbara, Sardinia, Italy 230421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola corsica 4 Limbara, Sardinia, Italy 230421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola corsica 5 Limbara, Sardinia, Italy 230421 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cucullata 1 Storrs, Connecticut 900508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola cucullata 2 Storrs, Connecticut 900508 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola declinata 2 Rodnea, Romania 090612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola declinata 3 Rodnea, Romania 090612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola declinata 4 Rodnea, Romania 090612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola delavayi Yunnan 1107 - Photo: Unknown (© 2014 Jan Thomas Johansson).

Viola doerfleri 1 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola doerfleri 2 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola doerfleri 4 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola doerfleri 5 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 1 Lenstad, Torslunda, Öland 120603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 11 Karum, Högsrum, Öland 150607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 12 Karum, Högsrum, Öland 150607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 15 Karum, Högsrum, Öland 150607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 2 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 3 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 4 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 6 Tveta, Torslunda, Öland 010609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola elatior 7 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola epipsila 1 Välleröd, Högestad, Skåne 810610 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola eximia 1 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola eximia 2 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola eximia 3 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola eximia 4 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola eximia 5 Mt Voras (Kajmakcalan), N Greece 180701 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola filicaulis 1 Pureora Forest Reserve NZ 151118 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola ganiatsasii 2 Partheno Dasos, Rhodopi Mountains, N Greece 180704 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola jooi 2 (cult) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola jordanii 2 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola kitaibeliana 3 above Seki, near Syedra, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola kitaibeliana 5 Nagykovácsi, Hungary 240504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola kitaibeliana 6 Nagykovácsi, Hungary 240504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola lanaiensis 4 Olinda Endangered Plant Facility, East Maui, Hawaii 130718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola lanaiensis 6 Olinda Endangered Plant Facility, East Maui, Hawaii 130718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola lanaiensis 7 Olinda Endangered Plant Facility, East Maui, Hawaii 130718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola lanaiensis 9 Olinda Endangered Plant Facility, East Maui, Hawaii 130718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 10 Röddinge Sk 140501 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 2 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola mirabilis 3 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola odorata 6 Lund Sk 140323 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmata 7 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmata 8 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 1 LP 4 to Roque Muchachos, La Palma 090430 (Richard Åkesson) - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 10 below the observatory, La Palma 070417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 11 below the observatory, La Palma 070417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 12 below the observatory, La Palma 070417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 2 LP 4 to Roque Muchachos, La Palma 090430 (Richard Åkesson) - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 5 below the observatory, La Palma 070417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 7 below the observatory, La Palma 070417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 8 below the observatory, La Palma 070417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palmensis 9 below the observatory, La Palma 070417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 3 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola palustris 6 Rövarekulan, Gudmuntorp Sk 210518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pedatifida 4 Lund Bot Garden 100527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pedunculata 1 Triumphal Park, CA 130410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pedunculata 2 Triumphal Park, CA 130410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pubescens 1 Storrs, Connecticut 900512 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pubescens 2 Storrs, Connecticut 900511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 10 Eketorp, Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 12 Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 13 Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 14 Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 15 Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 16 Gräsgård, Öland 150604 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 5 Eskilstorps Ängar, Skåne 850602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola pumila 6 Ekelunda, Sandby, Öland 120603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 4 Lund Bot Garden 080422 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola reichenbachiana 8 Röddinge Sk 140415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola riviniana ssp riviniana 2 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola rupestris 3 S of Vitemölla, Vitaby Sk 140419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola rupestris 5 S of Vitemölla, Vitaby Sk 140419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola rupestris 6 S of Vitemölla, Vitaby Sk 140419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola scorpiuroides 2 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola scorpiuroides 5 Vai, Crete 110330 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola selkirkii 1 Halsviksravinen, Ångermanland 870607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola selkirkii 3 Halsviksravinen, Ångermanland 870607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola sororia 4 Lund Bot Garden 150503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 10 South of Gustav Adolf Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 12 South of Gustav Adolf Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 13 Norra Lingenäset, Kristianstad Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 14 Norra Lingenäset, Kristianstad Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 15 Norra Lingenäset, Kristianstad Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 2 Lenstad, Torslunda, Öland 120603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 3 Norra Lingenäset, Kristianstad, Skåne 880608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 4 Norra Lingenäset, Kristianstad, Skåne 880608 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola stagnina 8 South of Gustav Adolf Sk 150628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola striata 1 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola striata 2 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola striata 7 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola striata 9 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola trichopetala 2 Dali, Yunnan 110706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola trichopetala 3 Dali, Yunnan 110706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola trichopetala 4 Dali, Yunnan 110706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp curtisii 4 Kämpinge, Räng Sk 080503 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp macedonica 1 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp macedonica 3 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp tricolor 4 Höör Sk 140713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp tricolor 5 Höör Sk 140713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp tricolor 6 Höör Sk 140713 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp tricolor 7 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola tricolor ssp tricolor 8 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola uliginosa 1 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola uliginosa 2 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola uliginosa 4 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola uliginosa 7 Anderstorp, Össjö Sk 140427 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Viola urophylla Baimashan, Yunnan 110710 - Photo: Unknown (© 2014 Jan Thomas Johansson).

Viola velutina 1 the Vigla Pass, Mt Varnous, N Greece 180630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).