Rhamnus 1a Santa Catalina Island, CA 130407 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus 1b Santa Catalina Island, CA 130407 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus 2b Santa Margarita River, CA 130415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus 2c Santa Margarita River, CA 130415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus 2d Santa Margarita River, CA 130415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 1 Ojén Andalusien SP 120319 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 11 Sierra Blanca de Ojén, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 13 Sierra Blanca de Ojén, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 2 Ojén Andalusien SP 120319 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 3 Ojén Andalusien SP 120319 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 4 Ojén Andalusien SP 120319 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 5 Albufera, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alpina 1 Copenhagen Bot Garden 140424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alpina 2 Copenhagen Bot Garden 140424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alpina 3 Copenhagen Bot Garden 140424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alpina 4 Copenhagen Bot Garden 140424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alpina 5 Copenhagen Bot Garden 140424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alpina 6 Copenhagen Bot Garden 140424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus cathartica 1 Lund Bot Garden 130528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus cathartica 2 Lund Bot Garden 130528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus cathartica 3 Lund Bot Garden 130528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus cathartica 4 Lund Bot Garden 130528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus cathartica 5 Lund Bot Garden 130528 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus cathartica 6 Lund Bot Garden 130906 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus costata 1 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus costata 2 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus costata 3 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus costata 4 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus costata 5 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus costata 6 Copenhagen Bot Garden 150626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus crenulata 1 Mirador de Don Martín, Tenerife 970320 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus crenulata 2 Mirador de Don Martín, Tenerife 970320 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus crenulata 4 Miradór de San Pedro, Tenerife 160404 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus crenulata 5 Miradór de San Pedro, Tenerife 160404 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus crenulata 6 Barr. del Infierno, Tenerife 160406 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus crenulata 7 Barr. del Infierno, Tenerife 160406 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus crocea 1 Rancho Santa Ana Botanic Garden, Los Angeles 130406 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus crocea 2 Rancho Santa Ana Botanic Garden, Los Angeles 130406 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus frangula 1 Lärkesholm, Tåssjö Sk 120829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus frangula 2 Lärkesholm, Tåssjö Sk 120829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus frangula 3 Lärkesholm, Tåssjö Sk 120829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus frangula 4 Lärkesholm, Tåssjö Sk 120829 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus frangula 6 Ljunghusen, Stora Hammar Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus glandulosa 1 Cubo de Galga, LP 090507 (Richard Åkesson) - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus glandulosa 2 Cubo de Galga, LP 090507 (Richard Åkesson) - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus glandulosa 3 La Galga, La Palma 070419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus glandulosa 4 La Galga, La Palma 070419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus globosa 1 Copenhagen Bot Garden 130913 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus globosa 2 Copenhagen Bot Garden 130913 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus imeretina 1 (cult) Skärva, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus imeretina 10 (cult) Skärva, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus imeretina 2 (cult) Skärva, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus imeretina 3 (cult) Skärva, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus imeretina 4 (cult) Skärva, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus imeretina 5 (cult) Skärva, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus imeretina 6 (cult) Skärva, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus imeretina 7 (cult) Skärva, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus imeretina 8 (cult) Skärva, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus imeretina 9 (cult) Skärva, Blekinge 160602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus integrifolia 3 La Fortaleza, Tenerife 090518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus integrifolia 4 La Fortaleza, Tenerife 090518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus integrifolia 5 La Fortaleza, Tenerife 090518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus integrifolia 6 La Fortaleza, Tenerife 090518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus ludovici-salvatoris 1 Soller Jardin Botanico, Mallorca 110419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus ludovici-salvatoris 2 Soller Jardin Botanico, Mallorca 110419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus lycioides 1 Desfiladero de Los Gaitanes, Andalucía Spain 220411 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus lycioides 2 Desfiladero de Los Gaitanes, Andalucía Spain 220411 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus lycioides 3 Desfiladero de Los Gaitanes, Andalucía Spain 220411 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus lycioides 4 Desfiladero de Los Gaitanes, Andalucía Spain 220411 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus myrtifolia 1 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus myrtifolia 2 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus myrtifolia 3 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus myrtifolia 4 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus myrtifolia 5 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus myrtifolia 6 Peñas Blancas, Sierra Bermeja, Andalucía Spain 220416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus oleoides 1 Zedeà, Armenia 180510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus oleoides 2 Zedeà, Armenia 180510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus oleoides 3 Zedeà, Armenia 180510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus oleoides 4 Zedeà, Armenia 180510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus pumila 1 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus pumila 2 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus pumila 3 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus pumila 4 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus rupestris 1 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus rupestris 2 Lund Bot Garden 130819 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus 2a Santa Margarita River, CA 130415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 10 Sierra Blanca de Ojén, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 12 Sierra Blanca de Ojén, Andalucía Spain 220414 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 6 Comboire FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 7 Comboire FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 8 Comboire FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus alaternus 9 Comboire FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus crocea 3 Rancho Santa Ana Botanic Garden, Los Angeles 130406 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus frangula 5 Ljunghusen, Stora Hammar Sk 130804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus integrifolia 1 La Fortaleza, Tenerife 090518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus integrifolia 2 La Fortaleza, Tenerife 090518 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Rhamnus pumila 5 between Villar d'Arène and Col du Lautaret FR 150707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).